Heterothops hosmanni ( Bernhauer, 1912 ) Bernhauer, 1912

Maus, Mariana Chani-Posse De, 2009, Revision of some types of Philonthus Stephens and Gabrius Stephens from southern South America (Coleoptera: Staphylinidae), with nomenclatural notes, Zootaxa 2034, pp. 31-42 : 34-35

Heterothops hosmanni ( Bernhauer, 1912) , comb. nov.

Philonthus hosmanni Bernhauer, 1912: 41 ; Bernhauer & Schubert, 1914: 341 (catalog); Bruch, 1915: 445 (catalog); Blackwelder, 1944: 133 (list); Herman, 2001: 2840 (catalog); Chani-Posse, 2004: 231 (list).

Type material. Holotype, Ƥ, “Rep. Argentina / Prov. Buenos Aires / Hosmann”, “ hosmanni Brh. / Typus ” (yellow label), “Chicago NHMus. / M. Bernhauer Collection”, “ Holotype Philonthus hosmanni Bernhauer, 1912 / Des. Chani-Posse de Maus 2008” ( FMNH).

Notes. In the original description Benhauer (1912) stated that he had a unique specimen of P. hosmanni . The specimen mentioned above agrees with the original description by Bernhauer (1912), being the holotype fixed by monotypy ( ICZN 1999, Article 73.1.2).

Comments. My examination of the holotype established that this species does not belong to the genus Philonthus (Philonthina) but belongs to the genus Heterothops Stephens, 1829 (Quediina) . In Smetana’s (1995) key to world subtribes of Staphylinini , this species will go to couplet 7, where it shares with Quediina “the inflexed lateral portion of anterior pronotal angle strongly inflexed, meeting prosternum at sharp” and “head with infraorbital ridge”. Based on the comparison of this species with the key to the Nearctic genera of Quediina by Newton et al. (2000), the diagnostic characters that define the North temperate genera of Quediina ( Smetana & Davies 2000), and other available keys and descriptions ( Lynch-Arribálzaga 1884; Sharp 1884; Bernhauer 1927; Coiffait & Sáiz 1966, 1968) for the genera of Quediina known from the Neotropics ( Herman 2001), Philonthus hosmanni shares with Heterothops the first antennal segment not longer than the next two combined, the last segment of maxillary palpus distinctly narrowed toward acute apex and shorter than the penultimate segment, and the nuchal ridge not distinctly converging with the infraorbital ridge.