Plusioglyphiulus similis, Golovatch & Geoffroy & Mauriès & Den Spiegel, 2009

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Den Spiegel, Didier Van, 2009, Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae), Zoosystema 31 (1), pp. 71-116 : 103-106

Plusioglyphiulus similis n. sp.

( Figs 27-30)

TYPE MATERIAL. — Indonesia. Borneo, KalimantanTimur, Pengadan , 01.257°N, 117.706°E, Cave Gua Ampanas , by hand, 18.VIII.2004, leg. A. Bedos & L. Deharveng (KAL-113), holotype ♂ ( MZB) ; 1 ♂, 1 ♀ paratypes ( MNHN GA062); 1 ♂ paratype ( SEM). — Same cave, 18.VIII.2004, leg. Y. R. Suhardjono & C. Rahmadi (KAL-YC04-09), 1 ♀ paratype ( MZB); 1 ♂, 1 ♀ paratypes ( ZMUM); 1 ♀ paratype ( ZMUC); 1 ♀ paratype ( VMNH).

ETYMOLOGY. — To emphasize the strong similarities to P. pallidior n. sp.

DIAGNOSIS. — Differs from the most similar P. pallidior n. sp. in the smaller size, the presence of three rows of bituberculate crests below the poriferous cones on most of the body segments and the somewhat longer legs.

DESCRIPTION

Length 17-27 mm, width 1.0- 1.6 mm, segments of posterior third of body being broadest.

Coloration marbled brown, sometimes with broad, paramedian, yellow stripes dorsally; head (except for blackish ocelli), collum and segment 2 mostly pale yellow; tips of metatergal tubercles usually brown, those of poriferous tubercles dark brown; antennae and legs light brown.

Body with 42-50p+3-1a+T. Length of holotype about 26 mm, width 1.6 mm, with 50p+1a+T.

Antennae rather short and clavate ( Figs 27A; 30A), antennomeres 6 and 7 each with a small group of bacilliform sensilla distodorsally.Ocellaria transversely subtriangular, with 8-11 ocelli in 5-7 longitudinal rows. Gnathochilarium slightly more densely setose due to a few smal setae on promentum ( Fig. 30B).

Postcollar constriction very evident, collum and segment 2 particularly strongly enlarged ( Fig. 27A, B). Carinotaxy formula of collum: 1p/t+2p/t+3p/t/t+// (ta)/(t)+4p/t/t +5p/t/t/t+//(ta)/(t)+pp/t/t/t+//ma/t ( Fig. 27B). Collum slightly depressed near anterior third. Carinotaxy of metatergum 2, 7/7+m/m+7/7; formula of metaterga 3 and 4, 4/4+3+M+3+4/4; formula of metaterga 5-7 and of several segments before apodous, 3/3+I/i+3/3+m/m+3/3+I/i+2/2; formula of midbody metaterga, 3/3+I/i+3/4(5)/3+m// m+3/4(5)/3+I/i+3/3; that of apodous segments, 4/4+m/m+4/4 ( Fig. 27 A-E); all crests and tubercles rather low, poriferous cones (I) considerably higher, but still broader than high, while median ones (m) as high as others ( Figs 27 A-E; 28A). Dorsal crests on several posteriormost segments considerably higher than others ( Fig. 27E). Midbody segments circular in cross-section ( Fig. 28A). Pleural regions of segments 2-4 conspicuously expanded, flap-shaped, especially so on segment 3 ( Fig. 27A). Pleural ridge behind gonopod aperture on male segment 7, tegument and limbus as in above congeners. Epiproct short, regularly and broadly rounded apically, with a paramedian pair of high but short crests/tubercles sometimes flanked on each side by a small knob at about midway; paraprocts more or less regularly convex, not flattened; hypoproct only slightly emarginate at caudal margin ( Fig. 27E, F).

Legs relatively long, on midbody segments about as long as body diameter ( Figs 28A; 30F). Claw at base with a short accessory spine ( Fig. 28B). Tarsi and their distal setae, as well as male legs 1-3 ( Figs 28C, D; 29A; 30 C-E) typical.

Both anterior and posterior gonopods ( Figs 29 B-D; 30G-I) very much like in P. pallidior n. sp., but both coxal processes cxp1 and cxp2, and arm (d), slightly different in shape.

REMARKS

Any obvious troglomorphic traits in this species seem to be absent.