Capobula, Haddad & Jin & Platnick & Booysen, 2021

Haddad, Charles R., Jin, Chi, Platnick, Norman I. & Booysen, Ruan, 2021, Capobula gen. nov., a new Afrotropical dark sac spider genus related to Orthobula Simon, 1897 (Araneae: Trachelidae), Zootaxa 4942 (1), pp. 41-71 : 52

publication ID	https://doi.org/ 10.11646/zootaxa.4942.1.2
publication LSID	lsid:zoobank.org:pub:79353662-7653-4F41-8B39-40E6E4B2E005
DOI	https://doi.org/10.5281/zenodo.4618497
persistent identifier	https://treatment.plazi.org/id/03D67155-FFA4-FFAD-FF25-FF2DA5BC2AF4
treatment provided by	Plazi
scientific name	Capobula
status	gen. nov.

Treatment

Key to the species of Capobula View in CoL gen. nov.

1 Females ............................................................................................. 2

– Males (males of C. neethlingi View in CoL spec. nov. and C. ukhahlamba View in CoL spec. nov. unknown).................................. 6

2 Abdomen grey, with cream inverted Y-shaped marking dorsally ( Fig. 10 View FIGURES 3–10 ); epigyne with large circular depressions incorporating copulatory openings ( Fig. 73 View FIGURES 71–74 )........................................................ C. ukhahlamba View in CoL spec. nov.

– Abdomen uniformly grey dorsally, without marking (e.g. Fig. 3 View FIGURES 3–10 ); epigyne with curved epigynal ridges incorporating copulatory openings............................................................................................ 3

3 Copulatory openings in anterolateral corners of faint M-shaped ridges ( Fig. 54 View FIGURES 54–58 ); bursae clearly much larger than primary spermathecae ( Fig. 55 View FIGURES 54–58 )................................................................... C. capensis View in CoL spec. nov.

– Copulatory openings in small C- or J-shaped ridges (e.g. Fig. 59 View FIGURES 59–64 ); bursae same size or smaller than primary spermathecae (e.g. Fig. 60 View FIGURES 59–64 )............................................................................................. 4

4 Anterior margin of ridges incorporating copulatory openings in same transverse plane as anterior margin of primary spermathecae; initial 2/3 of copulatory ducts before posterior bend heavily sclerotized and darker than last 1/3 ( Fig. 55 View FIGURES 54–58 )................................................................................................ C. infima View in CoL comb. nov.

– Anterior margin of primary spermathecae clearly in front of anterior margin of ridges incorporating copulatory openings; only initial 1/3 of copulatory ducts before posterior bend heavily sclerotized, last 2/3 clearly lighter in colour ( Figs 65 View FIGURES 65–70 , 71 View FIGURES 71–74 )...... 5

5 Ridges incorporating copulatory openings large and almost transverse to longitudinal plane of epigyne ( Fig. 71 View FIGURES 71–74 ); heads of bursae converging ( Fig. 72 View FIGURES 71–74 ) ( South Africa: Western Cape, Fig. 75 View FIGURE 75 )............................... C. neethlingi View in CoL spec. nov.

– Ridges incorporating copulatory openings small and obliquely orientated, at 45 degrees to longitudinal plane of epigyne ( Fig. 65 View FIGURES 65–70 ); heads of bursae diverging ( Fig. 66 View FIGURES 65–70 ) (eastern half of South Africa, Fig. 75 View FIGURE 75 ).................... C. montana View in CoL spec. nov.

6 Embolus as wide as 1/2 of tegulum width at midpoint, tip sharply curved and almost transverse to longitudinal plane of palp ( Fig. 62 View FIGURES 59–64 )................................................................................................................................................................................ C. infima View in CoL comb. nov.

– Embolus narrow, width less than ¼ of tegulum width at midpoint, tip directed disto-retrolaterally ( Figs 57 View FIGURES 54–58 , 68 View FIGURES 65–70 )…7

7 Embolus short, with stout tip ( Fig. 57 View FIGURES 54–58 ); femoral apophysis a sharp retrolateral spike ( Fig. 58 View FIGURES 54–58 ) ( South Africa: Western Cape, Fig. 75 View FIGURE 75 )................................................................................ C. capensis View in CoL spec. nov.

– Embolus longer, with slender tip ( Fig. 68 View FIGURES 65–70 ); femoral apophysis a stout ventral lobe ( Fig. 69 View FIGURES 65–70 ) (eastern half of South Africa, Fig. 75 View FIGURE 75 )................................................................................ C. montana View in CoL spec. nov.