Arcyphysa, Wells & Contents, 2018

Wells, Alice & Contents, Arturs Neboiss Table Of, 2018, Australian Diplectroninae reviewed (Insecta: Trichoptera), with description of 21 new species, most referred to a new genus, Zootaxa 4415 (1), pp. 1-44 : 24

Key to males of species of Arcyphysa

1 Apicolateral processes length at least 4x length of coxopodites of gonopods ( Figs 144–145 )....... Arcyphysa nebo sp. nov.

-. Apicolateral processes length less than 4x length of coxopodites................................................ 2

2(1) Gonopods with coxopodites in lateral view subquadrate to irregularly stout, with length equal to or less than 2x width (e.g., Figs 133, 137, 139 ).................................................................................... 3

-. Gonopods with coxopodites in lateral view more or less rectangular or rod-shaped, with length exceeding 2x maximum width (e.g., 105, 118, 129)................................................................................... 7

3 (2) In ventral view, apicolateral processes on segment IX strongly curved, distally club-shaped ( Fig. 134 ), coxopodites triangular; in lateral view, coxopodites sub-quadrilateral ( Fig. 135 )................................ Arcyphysa anaplasis sp. nov.

-. In ventral view, apicolateral processes tapered to acute apices or narrowly rounded apically, shape of coxopodites varied, not triangular in ventral view and sub-quadrangular in lateral view.................................................. 3

4(3) Harpagones each single, tapered ( Fig. 141 )........................................... Arcyphysa volsella sp. nov.

-. Harpagones each distinctly bilobed (e.g., Figs 132, 136 )....................................................... 5

5(4) Coxopodites of gonopods sub-rectangular in ventral and lateral views ( Figs 132–133 )........... Arcyphysa acmea sp. nov.

-. Coxopodites of gonopods triangular in ventral view ( Figs 136, 138 ).............................................. 6

6(5) Apicolateral processes on segment IX twisted, sinuous ( Figs 136–137 )....................... Arcyphysa tortula sp. nov.

-. Apicolateral processes on segment IX not as above ( Figs 138–139 )........................... Arcyphysa flinti sp. nov.

7(2) Phallic apparatus with parameres visible towards base; harpagones usually sharply and shallowly bifid apically (e.g., Figs 117, 122 )............................................................................................... 8

-. Phallic apparatus without parameres; harpagones never bifid apically, sometimes cup- or spatula-shaped (e.g., Figs 104 , 110)................................................................................................... 12

8(7) Ventral lobe of each apicolateral process on segment IX elaborately crescent-shaped in lateral view, with curves lined with spe- cialised setae ( Figs 118, 120 )........................................................................... 9

-. Ventral lobe of each apicolateral process not as above...................................................... 10

9(8) Apicolateral processes in lateral view bilobed, each with ventralmost lobe loosely crescent-shaped; tergite X covered with dense microtrichia ( Fig. 118 )....................................................... Arcyphysa caldera sp. nov.

Figures

FIGURES 132–145. Arcyphysa gen. nov. spp., ♂ genitalia diagnostic features. 132–133, Arc. acmea sp. nov. (Qld, Mt Bartle Frere, 0.5km N of South Peak [PT-999]): 132, ventral;133, left lateral.134–135, Arc.anaplasis sp. nov. (Qld, 25 km along Mt Lewis Road, SW of Mossman [PT-1003]): 134, ventral; 135, left lateral. 136–137, Arc. tortula sp. nov. (Qld, Bellenden Ker Range, Cable Tower No. 3 [PT-1031]): 136, ventral; 137, left lateral. 138–140, Arc. flinti sp. nov. (Qld, Palmerston National Park, Tchooratippa Creek [PT-2007]): 138, ventral; 139, left lateral; 140, dorsal. 141–142, Arc. volsella sp. nov. (Qld, Queensland, Lamington National Park, Coomera Creek [PT-1013]): 141, ventral; 142, left lateral.143–145, Arc. nebo sp. nov. (Qld, Mt Nebo [PT-1096]): 143, dorsal;144, left lateral; 145, ventral.

FIGURES 117–131. Arcyphysa gen. nov. spp., ♂ genitalia diagnostic features. 117–118, Arc. caldera sp. nov. (Qld, The Crater,Atherton [PT-1040]):117, ventral; 118, leftlateral.119–120, Arc.crescentinasp.nov. (Qld, TullyFalls [PT-1002]):119, ventral; 120, left lateral. 121–122, Arc.hugginsi sp. nov. (Qld, Bellenden Ker Range [PT-1000]): 121, ventral; 122, left lateral. 123–125, Arc. sybillae sp. nov. (Qld, Bloomfield River [PT-1016]): 123, ventral; 124, left lateral; 125, dorsal. 126–127, Arc. carnarvona sp. nov. (holotype; Qld, Carnarvon Gorge National Park): 126, ventral; 127, left lateral. 128–129, Arc. naumanni sp. nov. (holotype; Qld, 11 km W by N Bald Hill, McIllraith Range): 128, ventral; 129, left lateral. 130–131, Arc.lovedayi sp. nov. (holotype): 130, ventral; 131, left lateral.

FIGURES 102–109. Arcyphysa gen. nov. spp., diagnostic features.102–103, Arc.diamontona sp. nov., Vic., Diamond Creek, 7 km SE of Gembrook [PT-1006], ♂: 102, right fore- and hind wings, dorsal, 103, maxillary palp. 104–109, ♂ genitalia. 104– 105, Arc. angusta (Banks 1939; NSW, Styx R. [PT-1005]): 104, ventral, and 105, left lateral. 106–107, Arc. diamontona sp. nov. [PT-1006]:106, ventral, and 107, left lateral.108–109, Arc.fraserensis sp. nov. (Qld, Fraser Island, Pile Valley, nr Central Station [PT-1125]):108, ventral,and 109, leftlateral.Scale bars: Fig.102 = 1 mm; Fig.103 = 0.5 mm.Abbreviations:ac= anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; cox = coxopodite; fI, fII, fIII, fIV, fV = forks I–V; mc= median cell; har = harpago; ph = phallic apparatus; IX, X = abdominal segments IX, X