Odontomma patulа (Ponomarenko, 1985), Strelnikova & Yan, 2023

Strelnikova, O. D. & Yan, E. V., 2023, ON SPLITTING OF THE GENUS NOTOCUPES (COLEOPTERA: ARCHOSTEMATA): NEW DATA ON MORPHOLOGY AND TAXONOMY, Far Eastern Entomologist 488, pp. 1-28 : 23

Odontomma patulа (Ponomarenko, 1985) , comb. n.

Figs 9–12

Notocupes patulus Ponomarenko, 1985: 48 , tab. VI, fig. 1.

MATERIAL EXAMINED. Holotype, PIN no. 4034/2 – print and counterprint of elytron, metaventrite, and abdomen, apices of elytron and abdomen not preserved; Layma locality, Tyumen Formation , Middle Jurassic.

DIAGNOSIS. Abdomen covered with large, sparse tubercles, density 175 tb/mm 2. Elytron has up to around 38 small cells in the row.

DESCRIPTION. Epipleuron wide, gradually curving along its length, weakly narrowing at basal half to about 1/3 of its original width. Elytral base straight; humeral bulge distinct, rounded. Elytral cells small, more than 30 cells in a single row. Metaventrite trapezoid, his width 2.5 times exceeds length. Paracoxal suture do not reaches posterior angles of metaventrite. Central portion of abdominal sternites covered with large tubercles (0.05–0.07 mm in diameter; 0.02–0.06 distance between tubercles). Metacoxae strongly transverse, width two times exceeds length, covering half of first visible abdominal sternite.

MESUREMENS. Approximate body length 13, length of preserved body part 8.7, approximate elytral length 10, width at basal third 4.

COMPARISION. Similar to Odontomma trachylаena Ren et al., 2006 and О. sulcatum (Tan et al., 2007) in having small elytral cells and paracoxal suture not reaching posterior margins of metaventrite. Differs from O. trachylаena in noticeably smaller size and from О. sulcatum in smaller density of cuticular tubercles, and from both species in narrower epipleuron.

REMARKS. This species cannot be assigned to any of the five genera previously comprised former Notocupes . Here it is transferred from Notocupes to Odontomma Ren et al., 2006 on the basis of follow features: body covered with large, sparse tubercles; elytral base straight, humeral bulge distinct; epipleural margin dentate along its length; rather wide, only weakly narrowing posteriorly, epipleuron. There are no species with posteriorly widening epipleuron, if epipleuron is wide near elytral base it is usually strongly narrowing at the elytron midlength ( Figs 4 , 25–28 ).

Figures

Figs 9–12. Odontomma patula (Ponomarenko, 1985). 9 – photograph of the print; 10 – photograph of the counterprint; 11 – interpretative linedrawing of the print; 12 – interpretative linedrawing of the counterprint. Cuticular tubercles depicted in a square frame. Scale bar = 1 mm. 6

Figs 1–4. Types of elytral venation. 1 – Conexicoxa brachicephala (Ponomarenko, 1994); 2 – schematic drawing of Zygadenia spp.; 3 – Z. liui Jarzembowski et al., 2015; 4 – Z. westraliensis (Riek, 1968). (Figs 1, 2, 4 – redrawing with changes from Ponomarenko, 1994, 2006; Martin, 2010). Scale bar = 1.

Figs 25–28. Elytron venation types of five investigated genera of Archostemata. 25, 26 – Zygadenia alexrasnitsyni Strelnikova et Yan, 2021: 25 – photograph; 26 – interpretative linedrawing; 27, 28 – Notocupes elegans Ponomarenko, 1994: 27 – photograph; 28 – interpretative linedrawing. Elytral fields marked with Roman numerals, veins – with Arabic. Scale bar = 1 mm.