Haroldiellus lansbergei (Harold)

Skelley, Paul E. & Keller, Oliver, 2022, A third species of Haroldiellus Gordon and Skelley, 2007 from Mexico and Central America (Coleoptera: Scarabaeidae: Aphodiinae: Aphodiini), Insecta Mundi 2022 (916), pp. 1-11 : 3

Haroldiellus lansbergei (Harold)

Figures 2 , 7 , 10

Aphodius lansbergei Harold 1874: 179 ~ Bates 1887: 84; Blackwelder 1944: 212; Dellacasa et al. 1998: 158 (habitus, epipharynx and aedeagus figured; lectotype designated).

Aphodius (Bodilus) lansbergei Harold ~ Schmidt 1913: 123.

Agrilinus lansbergei (Harold) ~ Dellacasa et al. 2002: 161.

Haroldiellus lansbergei (Harold) ~ Gordon and Skelley 2007: 270.

Diagnosis. Length 4.3–5.0 mm, width 1.9–2.4 mm. A member of Haroldiellus distinguished from other species by: Elytral striae weakly impressed, intervals weakly convex, interval color pale, lacking distinct markings ( Fig. 2 ). Female lacking median frontal tubercle; male median frontal tubercle transverse ( Fig. 7 ). Protibia with dorsal surface lacking punctures.

Distribution. ( Fig. 10 ). Mexico (Chiapas, Hidalgo, Michoacán, Oaxaca, Querétaro, San Luis Potosí, Veracruz). Records here are repeated from Dellacasa et al. 2002; Skelley et al. 2007; Halffter et al. 2012; Minor 2017.

Material examined. MEXICO: Chiapas / 10 km. e. LaTrinitaria/ August 14, 1967 / under dung/ W.W. Gibson // TAMU-ENTO/ X0081443/ Barcode // Haroldiellus / lansbergei / det.P.E.Skelley 2021 (1 TAMU); MEXICO: State of/ Hidalgo, 1Km.N / Jacala, 5000Ft. // R.E.Woodruff/ 24-X-70/ Cow Dung (4 FSCA); MEXICO: Hidalgo / Puerto de la Zora/ 24-X-1970, light/ R. E.Woodruff (1 FSCA); E. DO MICHOACAN / rd. Méx-37 Uruapan / – Nueva Italia / La Gallina dint. // m 1250- 23.07.2001 / N19°19′ -W102°04′ / M. Dellacasa & / I. Martinez leg. (1 CMN); MEXICO: Querétaro; Mpio:/ Jalpan; nr.Valle Verde,/ Bio.St.El Pilon/ 21°27′41″N, 99°10′41″W / 11-DEC-2013, 1284m/ P.Skelley, P.Kovarik, R. Jones (1 FSCA); Rancho Pextlán/ (Xico)/ 19°25′28.9″N / 97°01′43.0W / 19.III.2005 - m 1465 // MEXICO / Edo. Veracruz / Leg. González O. &/ Suarez T./ excr. bovino // Haroldiellus / lansbergei/ ( Harold, 1874)/ Det. Dellacasa, 2011 (1 FSCA); Rancho Pextlan/ Mpio. Xico/ 19°25′28.9″N / 97°01′43.0W / 12.IX.2005 - m 1465 // MEXICO / Edo. Veracruz / Leg. Cruz M. Cabrero F. & Suarez T./ excr. vaca // Haroldiellus / lansbergei/ ( Harold, 1874)/ Det. Dellacasa, 2011 (1 FSCA).

Remarks. The specimen from Querétaro is slightly larger than other specimens presently available. It also has the discal intervals of the elytra darkened, appearing as a single dark mark. Other morphological features like the more flattened intervals and lack of protibial punctures indicate it is H. lansbergei as presently understood. The Chiapas ( Mexico) record published in Minor (2017) represents most likely the new species described herein, but needs to be re-examined based on its locality.

Figures

Figure 10. Distribution map of Haroldiellus lansbergei (blue) as currently known from specimen labels and literature records. Chiapas, Mexico is striped because it is a literature record that may represent the new species H. woodruffi described herein. Map made with SimpleMappr (Shorthouse 2010).

Figures 1–5. Haroldiellus species. 1) H. sallei dorsal habitus. 2) H. lansbergei dorsal habitus. 3) H. woodruffi, holotype, dorsal habitus. 4) H. sallei protibia dorsal surface with punctures. 5) H. woodruffi protibia dorsal surface without punctures. Scale line = 0.5 mm.

Figures 6–9. Heads of Haroldiellus species. 6) H. sallei. 7) H. lansbergei. 8) H. woodruffi female allotype. 9) H. woodruffi male holotype. Scale line = 0.5 mm (all figures).