Dissomphalus, Ashmead, 1893

Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco & Azevedo, Celso O., 2018, New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil, Zootaxa 4462 (1), pp. 1-40: 4-6

publication ID

https://doi.org/10.11646/zootaxa.4462.1.1

publication LSID

lsid:zoobank.org:pub:FE62FF43-119A-4EBC-92FE-5012FFAB603E

persistent identifier

http://treatment.plazi.org/id/03D487B8-C47E-CD44-FF22-FD5E13E5FEE3

treatment provided by

Plazi

scientific name

Dissomphalus
status

 

Key to Dissomphalus   males of the Cerrado, Caatinga and Northern relicts of the Atlantic Forest , Brazil  

1. Tergal process absent ( amplus   species-group)............................................................... 2

- Tergal process present ( Figs. 13 View FIGURES 5–16 , 18, 22 View FIGURES 17–29 ).................................................................... 3

2. Mandible bidentate; posterior hypopygeal margin straight; aedeagal ventral ramus shorter than dorsal body ( Figs 5–8 View FIGURES 5–16 ).......................................................................... D. boitata Colombo & Azevedo   sp. nov.

- Mandible tridentate; posterior hypopygeal margin weakly concave; aedeagal ventral ramus longer than dorsal body ( Figs 9– 11 View FIGURES 5–16 )................................................................... D. boto Colombo & Azevedo   sp. nov.

3. Tergal process located medially ( Fig. 61 View FIGURES 56–70 )................................................................... 4

- Tergal process located laterally ( Figs. 13 View FIGURES 5–16 , 18, 22 View FIGURES 17–29 )............................................................. 9

4. Tergal process with one pair of conspicuous depressions, transversally elliptical, tubercle present, directed each other (vallensis species-group) ( Fig. 61 View FIGURES 56–70 )................................................................................ 5

- Tergal process with one pair of conspicuous depressions, never transversally elliptical, tubercle present, never directed each other................................................................................................ 6

5. Parameral apex convex and oblique; aedeagal ventral ramus as long as aedeagal dorsal body; inner lobe of aedeagal dorsal body never setose............................................................... D. bifurcatus Azevedo, 1999  

- Parameral apex never convex and oblique; aedeagal ventral ramus shorter than aedeagal dorsal body; inner lobe of aedeagal dorsal body setose ( Figs 60–65 View FIGURES 56–70 )........................................... D. torem Colombo & Azevedo   sp. nov.

6. Tufts of tergal process linear and horizontal ( linearis   species-group)......................... D. linearis Azevedo, 1999  

- Tufts of tergal process neither linear nor horizontal........................................................... 7

7. Tergal process with large tubercles; apex of aedeagal dorsal body concave; apex of aedeagal ventral ramus with row of small ventral teeth (longiclypeus species-group)............................................ D. scamatus Azevedo, 1999  

- Tergal process without tubercles; apex of aedeagal dorsal body never concave; apex of aedeagal ventral ramus without row of small ventral teeth..................................................................................... 8

8. Tergal process with depression shallow, tufts large; aedeagal dorsal body nearly as long as ventral ramus, apex with two large teeth (apertus species-group)..................................................... D. subpilosus Azevedo, 1999  

- Tergal process with depression deep; aedeagal dorsal body much longer than ventral ramus, apex without two large teeth (altivolans species-group)............................................................ D. megomphalus Evans, 1969  

9. Aedeagal ventral ramus very longer than dorsal body, tubular, wide, rounded apex, without apical filament (brasiliensis species-group).......................................................... D. tubulatus Redighieri & Azevedo, 2006  

- Aedeagal ventral ramus never tubular ( Figs. 15 View FIGURES 5–16 , 20, 24 View FIGURES 17–29 )...................................................... 10

10. Aedeagal ventral ramus with apical filament ( microstictus   species-group)................... D. microstictus Evans, 1969  

- Aedeagal ventral ramus without apical filament ( Figs. 15 View FIGURES 5–16 , 20, 24 View FIGURES 17–29 ).............................................. 11

11. Tergal process with callus; volsella with small basal projection ( Figs 80–84 View FIGURES 71–84 )....................................................................................................... D. matintaperera Colombo & Azevedo   sp. nov.

- Tergal process without callus; volsella without such projection................................................. 12

12. Tergal process with one pair of conspicuous depressions, tubercle present, pit on top, lateral margin setose ( Figs 71–75 View FIGURES 71–84 )......................................................................... D. caipora Colombo & Azevedo   sp. nov.

- Tergal process not as above............................................................................. 13

13. Tergal process with depression deeply excavated, depressions partially covered by flap........................................................................................................ D. ubracus Brito & Azevedo, 2017  

- Tergal process not deeply excavated, depressions not covered by flap........................................... 14

- Pits of tergal process never drop-shaped, usually circular..................................................... 15

15. Tuft of tergal process with one distinct long seta ( gilvipes   species-group) ( Fig. 27 View FIGURES 17–29 )................................. 16

- Tuft of tergal process without such seta................................................................... 18

16. Median clypeal lobe duckbill-shaped; gena with one pair spines ( Figs 25–29 View FIGURES 17–29 )........ D. irupe Colombo & Azevedo   sp. nov.

- Median clypeal lobe not duckbill-shaped; gena without spines................................................. 17

17. Notauli complete; apex of dorsal body narrowed in lateral view............................... D. gilvipes Evans, 1979  

- Notauli incomplete or absent; apex of dorsal body wide in lateral view..................... D. krombeini Azevedo, 1999  

18. Depressions of tergal process inclined and linear (rufipalpis species-group)...................... D. infissus Evans, 1969  

- Tergal processes without depressions or if present never inclined and linear....................................... 19

19. Depressions of tergal process ovoid, deep and large, about 0.5 × as long as tergite, rim usually setose (bicavatus species-group)................................................................................................... 20

- Depressions of tergal process not as above................................................................. 22

20. Aedeagal basal process without median tooth........................................ D. geniculatus Azevedo, 2001  

- Aedeagal basal process with median tooth................................................................ 21

21. Aedeagal basal process wide.................................................... D. lobisserratus Azevedo, 2001  

- Aedeagal basal process narrow and sharpened....................................... D. curvilongus Azevedo, 2001  

22. Tubercles of tergal process large with high rim and tuft densely setose ( conicus   species-group) ( Figs. 13 View FIGURES 5–16 , 18, 22 View FIGURES 17–29 )......... 23

- Tubercles of tergal process small........................................................................ 29

23. Basivolsela with thick, conspicuous bristles................................ D. laminaris Redighieri & Azevedo, 2004  

- Basivolsela without such bristles....................................................................... 24

24. Outer lobe of aedeagal dorsal body with warts............................. D. verrucosus Redighieri & Azevedo, 2004  

- Outer lobe of aedeagal dorsal body without warts.......................................................... 25

25. Aedeagal ventral ramus with apical half translucent, narrow, apex with two horn-shaped expansions................................................................................................... D. cervoides Azevedo, 2003  

- Aedeagal ventral ramus without such expansions............................................................ 26

26. Basivolsella with large, wide, rounded expansion directed upward at inner margin; ventral margin of aedeagal ventral ramus with deep median emargination; dorsal margin of outer lobe of aedeagal dorsal body strongly conical in lateral view........................................................................................ D. conicus Azevedo, 2003  

- Basivolsella without such expansion; aedeagal ventral ramus without emargination; dorsal margin of outer lobe of aedeagal dorsal body not conical in lateral view.................................................................... 27

27. Parameral apex bifurcated; aedeagal ventral ramus longer than dorsal body, with small basal extension ( Figs 12–16 View FIGURES 5–16 )........................................................................... D. curupira Colombo & Azevedo   sp. nov.

- Parameral apex not bifurcated; aedeagal ventral ramus equal or shorter than dorsal body, without basal extension......... 28

28. Mandible with three teeth; parameral apex serrated; aedeagal ventral ramus as long as the dorsal body ( Figs 21–24 View FIGURES 17–29 )............................................................................... D. saci Colombo & Azevedo   sp. nov.

- Mandible with two teeth; parameral apex rounded; aedeagal ventral ramus shorter than dorsal body ( Figs 17–20 View FIGURES 17–29 ).................................................................................. D. iara Colombo & Azevedo   sp. nov.

29. Depression of tergal process with edge without thick and conspicuous bristles (punctatus species-group) ( Figs. 31, 35, 39 View FIGURES 30–41 )...................................................................................................... 30

- Depression of tergal process with edge with thick and conspicuous bristles....................................... 33

30. Median clypeal lobe trapezoidal; inner margin of basivolsella with large rounded callus.............. D. napo Evans, 1979  

- Median clypeal lobe tridentate; inner margin of basivolsella without callus....................................... 31

31. Mandible with two teeth; parameral apex straight ( Figs 38–41 View FIGURES 30–41 )................ D. jurupari Colombo & Azevedo   sp. nov.

- Mandible with three teeth; parameral apex rounded.......................................................... 32

32. Posterior hypopygeal margin weakly concave; parameral apex bifurcated in ventral view ( Figs 30–33 View FIGURES 30–41 )..................................................................................... D. acutipupu Colombo & Azevedo   sp. nov.

- Posterior hypopygeal margin straight; parameral apex not bifurcated in ventral view ( Figs 34–37 View FIGURES 30–41 )....... D. cuca Colombo & Azevedo   sp. nov.

33. Tubercles of tergal process slightly directed mesad (tuberculatus species-group)............... D. plaumanni Evans, 1964  

- Tubercles of tergal process never directed mesad............................................................ 34

34. Paramere never peanut-shaped or S-shaped (setosus species-group)............................................. 35

- Paramere peanut-shaped or S-shaped ( Figs. 68, 69 View FIGURES 56–70 ).......................................................... 39

35. Dorsal margin of paramere with apical spine; aedeagal ventral ramus with apical half divided into two long tubular rami.......................................................................... D. magnus Redighieri & Azevedo, 2006  

- Dorsal margin of paramere without spine; aedeagal ventral ramus without such rami............................... 36

36. Mandible with four teeth............................................................................... 37

- Mandible with two or three teeth........................................................................ 38

37. Median clypeal lobe trapezoidal, median tooth straight, higher in lateral view; outer lobe of aedeagal dorsal body leaf-shaped ( Figs 47–50 View FIGURES 42–55 )........................................................ D. capelobo Colombo & Azevedo   sp. nov.

- Median clypeal lobe tridentate, median tooth rounded, with posterior and lateral sides high and lower anterior and central regions; outer lobe of aedeagal dorsal body never leaf-shaped ( Figs 51–55 View FIGURES 42–55 )....... D. iracema Colombo & Azevedo   sp. nov.

38. Mandible two teeth; median clypeal lobe undefined; posterior hypopygeal margin concave ( Figs 42–46 View FIGURES 42–55 ).................................................................................... D. boibumba Colombo & Azevedo   sp. nov.

- Mandible with three teeth; median clypeal lobe trapezoidal; posterior hypopygeal margin straight ( Figs 56–59 View FIGURES 56–70 ).............................................................................. D. joaquinae Colombo & Azevedo   sp. nov. 39. Median clypeal lobe trilobate; posterior hypopygeal margin weakly concave; paramere peanut-shaped ( Figs 66–70 View FIGURES 56–70 )............................................................................ D. boiuna Colombo & Azevedo   sp. nov.

- Median clypeal lobe undefined; posterior hypopygeal margin straight; paramere S-shaped ( Figs 76–79 View FIGURES 71–84 )................................................................................... D. mapinguari Colombo & Azevedo   sp. nov.