Scatimus, Erichson, 1847

Martínez-Revelo, Diego Esteban, Lopera-Toro, Alejandro & Medina, Claudia A., 2020, Review of Scatimus Erichson (Coleoptera: Scarabaeidae) in Colombia with the description of a new species, Zootaxa 4890 (4), pp. 521-534 : 531

Key to Scatimus species in Colombia based on Génier & Kohlmann (2003)

1. Carina of the head with lateral extremities posteriorly directed (62, Figs. 2E, F ) ( Scatimus ovatus clade)................ 2

- Carina of the head with lateral extremities anteriorly directed (63, Figs. 2 A – D ) ( Scatimus strandi clade)................ 3

2(1). Dorsal portion of the eye approximately twice as long as wide (80, Fig. 2E ), coarse setiferous punctures of pronotum present in lateral fossae and anteriorly to fossae (131, Figs. 3A, B ), apex of parameres swollen, more or less spherical ( Figs. 4B, F )....................................................................................... Scatimus fernandezi

- Dorsal portion of the eye variable in size (82, Fig. 2F ), coarse setiferous punctures of pronotum absent or variably distributed (133), apex of parameres flat and tapering into a point ( Figs. 4A, E )................................ Scatimus ovatus

Figures

FIGURE 2. Scatimus heads in dorsal view. A, Scatimus strenua male (Antioquia, 5°52’45.05’’N 75°12’2.33’’W); B, Scatimus strenua female (Antioquia, 5°53’16.68’’N 75°11’37.11’’W); C, Scatimus erinnyos male (Costa Rica, Parque Nacional Guanacaste); D, Scatimus strandi male (Nariño, 0°30’N 77°13’W); E, Scatimus fernandezi (Casanare, 5°16’20.9”N 72°24’2.6”W); F, Scatimus ovatus (Tolima, 05°08’58.3’’N 74°47’06.1’’W). Scale bars = 0.5 mm.

FIGURE 3. Scatimus pronotum, mesepimeron, metasternum, abdomen, and metafemur. A, Scatimus strandi pronotum (lateral view); B, Scatimus strenua pronotum (lateral view); C, Scatimus strandi mesepimeron (ventral view); D, Scatimus strenua mesepimeron (ventral view); E, Scatimus strenua metasternum (ventral view); F, Scatimus strenua abdomen; G, Scatimus erinnyos metafemur (ventral view); H, Scatimus strenua metafemur (ventral view). Scale bar = 0.5 mm.

FIGURE 4. Male genitalia of Scatimus from Colombia. A, Scatimus fernandezi aedeagus (lateral view, scale bar = 0.5 mm); B, Scatimus ovatus aedeagus (lateral view, scale bar = 0.5 mm); C, Scatimus strandi aedeagus (lateral view, scale bar = 0.5 mm); D, Scatimus strenua aedeagus (lateral view, scale bar = 0.5 mm); E, Scatimus fernandezi parameres (ventral view, scale bar = 0.25 mm); F, Scatimus ovatus parameres (ventral view, scale bar = 0.25 mm); G, Scatimus strandi parameres (ventral view, scale bar = 0.25 mm); H, Scatimus strenua parameres (ventral view, scale bar = 0.25 mm); I, Scatimus fernandezi superior right peripheral sclerite (SRP, scale = 0.25 mm); J, Scatimus ovatus superior right peripheral sclerite (SRP, scale = 0.25 mm); K, Scatimus strandi superior right peripheral sclerite (SRP, scale = 0.25 mm); L, Scatimus strenua superior right peripheral sclerite (SRP, scale = 0.25 mm); M, Scatimus fernandezi additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm; N, Scatimus ovatus additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm; O, Scatimus strandi additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm; P, Scatimus strenua additional (AS), subaxial (SA), and axial (A) sclerites superposed, scale bar = 0.25 mm.