Haroldiataenius, Chalumeau, 1981

Stebnicka, Z. Teresa & Skelley, Paul E., 2009, A revision of the genus Haroldiataenius Chalumeau (Scarabaeidae: Aphodiinae: Eupariini), Insecta Mundi 2009 (62), pp. 1-16 : 2-4

Key to Species of Haroldiataenius

1. Body psammodiiform (Fig. 12); lateral margins of pronotum and elytron with long setal fringe, setal length much greater than distance between setal bases ( Fig. 1 ). Clypeus transversely rugose. Mexico, Panama........................................................................ H. mariarum (Bates)

— Body psammodiiform to robustly elongate (Fig. 13-15); lateral margins of pronotum or elytron bare, or with fringe of short setae; if setae present, then length equal to or less than distance between setal bases ( Fig. 2, 5-11 ). Clypeus punctate, smooth, or weakly rugose ..................... 2

2(1). Meso- and metafemoral posterior margin with distinct, complete line from trochanter to apex ( Fig. 3 ) ....................................................................................................................................... 3

— Meso- and metafemoral posterior margin lacking complete line, if present, then only in distal half ( Fig. 4 )........................................................................................................................................ 4

3(2). Clypeus rounded each side of median emargination. Base of head lacking band of coarse punctures. Elytral intervals convex ( Fig. 5 ), lacking setae on disc, some setae may be visible on apical declivity. Florida, USA................................................................. H. saramari (Cartwright)

— Clypeus angulate to distinctly toothed each side of median emargination. Base of head with band of coarse punctures. Elytral intervals tectiform, with short setae ( Fig. 6 ). Baja California, Mexico and Arizona, USA....................................................................................... H. lucanus (Horn)

4(2). Lateral pronotal margin lacking distinct setae; lateral bead of pronotum gradually thickened toward anterior angle ( Fig. 7 ). Mexico, Guatemala, Honduras, El Salvador............................. .................................................................................................................. H. limbatus (Bates)

— Lateral pronotal margin with distinct setal fringe; lateral bead of pronotum fine throughout length, not distinctly thickened near anterior angles ............................................................... 5

5(4). Pronotal punctures of uniform size, punctures dense, separated by less than one diameter over entire surface ( Fig. 2 , 15); base of pronotum with marginal line obscured by coarse punctures. Mexico, Guatemala, Honduras.................................................................. H. hintoni (Saylor)

— Pronotal punctures mixed fine and coarse; coarse pronotal punctures more widely scattered, separated by at least their own diameter ( Fig. 8-11 ); base of pronotum with distinct marginal line ............ 6

6(5). Clypeus strongly angulate to dentate each side of median emargination. Elytral intervals with long scattered setae, most prominent laterally ( Fig. 8 ). Texas and Arizona, USA; Mexico....... ...................................................................................................... H. semipilosus (Van Dyke)

— Clypeus rounded or angulate each side of median emargination. Elytral intervals lacking setae ( Fig. 9-11 ). Southern Texas, USA............................................................................................. 7

7(6). Lateral elytral margin with indistinct fringe of setae, if visible, setae distinctly shorter than distance between their bases ( Fig. 9 ). Coarse pronotal punctures more widely scattered, diameter equal to or less than width of basal pronotal margin ...................... H. griffini (Cartwright)

— Lateral elytral margin with distinct fringe of short setae, setae as long as distance between their bases ( Fig. 10-11 ). Coarse pronotal punctures evenly distributed basally, their diameter generally greater than width of broad basal pronotal margin .................................................................. 8

8(7). Clypeal anterior margin evenly rounded each side of median emargination ( Fig. 10 ). Elytral intervals weakly convex on disc. Body dark brown, nearly black. Big Bend region, Texas, USA............ ...................................................................................................................... H. buvexus , n. sp.

— Clypeal anterior margin angulate each side of median emargination ( Fig. 11 ). Elytral intervals strongly convex on disc. Body dark reddish brown. Southern coastal Texas, USA................... .......................................................................................................... H. convexus (Robinson)

Figures

Figures 1-7. Haroldiataenius spp. 1) H. mariarum, lateral habitus. 2) H. hintoni lateral habitus. 3) H. lucanus, mesotibia. 4) H. semipilosus, mesotibia. 5) H. saramari, lateral habitus. 6) H. lucanus, lateral habitus. 7) H. limbatus, lateral habitus.

Figures 8-11. Haroldiataenius spp., lateral habitus. 8) H. semipilosus. 9) H. griffini. 10) H. buvexus, paratype. 11) H. convexus, holotype.