Anemesia infuscata, Zonstein, 2018

Zonstein, Sergei, 2018, A revision of the spider genus Anemesia (Araneae, Cyrtaucheniidae), European Journal of Taxonomy 485, pp. 1-100 : 44-47

publication ID	https://doi.org/ 10.5852/ejt.2018.485
publication LSID	lsid:zoobank.org:pub:55A0F74D-FA80-4C6A-AD74-B49C9061A449
DOI	https://doi.org/10.5281/zenodo.3848285
persistent identifier	https://treatment.plazi.org/id/84CDD144-26EB-4DFF-8BE2-F878AA5C7471
taxon LSID	lsid:zoobank.org:act:84CDD144-26EB-4DFF-8BE2-F878AA5C7471
treatment provided by	Valdenar
scientific name	Anemesia infuscata
status	sp. nov.

Treatment

Anemesia infuscata View in CoL sp. nov.

urn:lsid:zoobank.org:act:84CDD144-26EB-4DFF-8BE2-F878AA5C7471 Figs 14 View Figs 10–17 , 26 View Figs 18–26 , 38–41 View Figs 34–41 , 45–47 View Figs 42–47 , 61–62 View Figs 48–62 , 76–77 View Figs 63–77 , 87–89 View Figs 78–89 , 103 View Figs 99–107 , 115–116 View Figs 108–116 , 131–132 View Figs 125–132 , 186–187 View Figs 173–187 , 220–223 View Figs 215–223 , 258– 265 View Figs 245–265 , 289–290 View Figs 278–290 , 324–329, 346–353, 365–367, 373 View Fig

Diagnosis

Anemesia infuscata sp. nov. differs from A. incana in a considerably narrower clypeus and noticeably larger eyes both in males and females ( Figs 61–62 View Figs 48–62 , 76–77 View Figs 63–77 ; cf. Figs 59 View Figs 48–62 , 73 View Figs 63–77 ). From the most similarlooking A. infumata sp. nov., males of A. infuscata sp. nov. differ by a less tapering embolus which arises closer to the prolateral side of the palpal organ ( Figs 220–223 View Figs 215–223 ; cf. Figs 217–219 View Figs 215–223 ), while the conspecific females differ in their longer cone spermathecae ending with certainly dilated receptacles ( Figs 258–265 View Figs 245–265 ; cf. Figs 255–257 View Figs 245–265 ).

Etymology

The specific epithet is derived from the Latin adjective ‘ infuscatus ’, meaning ‘infuscate’, referring to the entirely darkened (blackened) colouration of these spiders. The gender is feminine.

Material examined

Holotype

TAJIKISTAN: Ƌ, foothills of Hissar Mts, Varzob Canyon, 7 km N of Dushanbe, vicinity of Harangon , 1060 m, 38°40′04′′ N, 68°46′56′′ E, 27 May 2002, S. Zonstein leg. ( SMNH).

GoogleMaps

Paratypes (19 ƋƋ, 37 ♀♀)

TAJIKISTAN: 1 Ƌ, 2 ♀♀, same collecting data as for the holotype ( SMNH); 1 ♀, same area, environs of Harangon 5 km N of Dushanbe, 1100 m, 38°39.9′ N, 68°47.7′ E, 3 May 2015, S. Zonstein leg. ( SMNH); 2 ƋƋ, 6 ♀♀, Sanglok Mts, 3–5 km W Sebiston, 1300–2200 m (38°15′ N, 69°17′ E), 2 May 1990, S. Zonstein leg. ( SMNH); 3 ƋƋ, 2 ♀♀, same collection data as for preceding but 7 May 1991, S. Ovchinnikov and S. Zonstein leg. ( SMNH); 2 ƋƋ, 5 ♀♀, same collection data as for preceding but 16–18 May 2002, S. Zonstein leg. ( SMNH); 4 ♀♀, same collection data as for preceding but 29 Apr. 2015, S. Zonstein leg. ( SMNH); 6 ƋƋ, 8 ♀♀, Hazratisho Mts, Sangdara Gorge, 1650 m, 38°22′ N, 70°09′ E, 19–25 May 2002, S. Zonstein leg. ( SMNH); 5 ƋƋ, same area, Iokunj Gorge, 1600–2000 m, 38°23′ N, 70°10′ E, 23–26 May 2002, S. Zonstein leg. ( SMNH); 3 ♀♀, Kuhitek Mts, 7 km NE of Hovaling, 1850 m, 38°21.6′ N, 70°03.9′ E, 28 Apr. 2015, S. Zonstein leg. ( SMNH, ZMMU); 3 ♀♀, Mazar Mts, Sultanmazar 16 km NNE of Hovaling, 1850 m, 38°28.3′ N, 70°04.1′ E, 28 Apr. 2015, S. Zonstein leg. ( SMNH); 3 ♀♀, same area, Darai-Mukhtor 5 km NW of Hovaling, 1600 m, 38°23.6′ N, 69°57.9′ E, 28 Apr. 2015, S. Zonstein leg. ( SMNH).

Other material examined (8 ♀♀, 2 ♀♀ subad.)

TAJIKISTAN: 1 ♀, Rangentau Mts, Fahrabad Pass, 1200–1300 m, 38°20′ N, 68°43′ E, 3 Apr. 1967, E. Andreeva leg. ( MIZW); 1 ♀ subad., Hazratisho Mts, 25 km E of Muminabad, 24 May 1966, E.Andreeva leg. ( MIZW); 1 ♀ subad., same collection data as for preceding but 13 km NE of Muminabad, 13 Jun. 1966, E. Andreeva leg. ( MIZW); 7 ♀♀, Kuhitek Mts, 1 km E of Hovaling, 1600–1700 m, 38°20′ N, 69°59′ E, 11 Oct. 1987, S. Zonstein and S. Ovchinnikov leg. ( SMNH, ZMMU).

Description

Male (holotype)

HABITUS. See Fig. 14. View Figs 10–17

MEASUREMENTS. TBL 17.80, CL 7.17, CW 6.28, LL 0.80, LW 1.24, SL 3.63, SW 3.33.

COLOUR. Carapace, palps and legs predominantly intensive dark reddish brown; anterior edge and lateral margins of carapace, chelicerae and femora even darker; whole eye tubercle brownish black; sternum, labium, maxillae and leg coxae light coffee brownish coloured, metatarsi and tarsi III–IV brownish orange; abdomen yellowish brown with dark chocolate brown pattern consisting of very broad median longitudinal stripe crossed medially and posteriorly with six wide procurved fasciae, and numerous small and partially fused spots located between them; ventral abdominal surface with darker horse-shoe spot covering genital area; book-lungs paler, light yellowish brown; spinnerets even paler, light yellow.

PROSOMA. Clypeus and eye tubercle as in Fig. 61 View Figs 48–62 . Eye diameters and interdistances: AME 0.20(0.29), 0.30, 0.27, 0.22, AME–AME 0.23(0.13), ALE–AME 0.23(0.19), ALE–PLE 0.14, PLE–PME 0.02, PME–PME 0.49. Cheliceral rastellum composed of 15–20 stout bristles lacking tips and located in front of fang base and on low mound. Cheliceral furrow with 6 promarginal teeth and 9–10 smaller retromarginal teeth each. Sternum, labium and maxillae as shown in Fig. 103 View Figs 99–107 . Sternal sigilla small,

posterior pair broadly oval and located distantly from sternal margin. Each maxilla with 11–12 cuspules confined to probasal maxillary edge.

SPINATION. Palp: femur d3, pd1; patella p1; tibia d1, p3, r1, v1+ca 20 stout bristles; tarsus d5–6. Leg I: femur d4, pd2; tibia p2, v6–7+m; metatarsus v5. Leg II: femur d4, pd2, rd1; patella p1; tibia p3, v8–9; metatarsus p1, v8. Leg III: femur d4, pd3, rd3–4; patella p2, r1; tibia d1, p3, r3, v7; metatarsus pd5, p5, r3, v9–11; tarsus p3. Leg IV: femur d4, p3, r3; patella r1; tibia r3, v9; metatarsus pd1, p2, r3, v13–15; tarsus pv9–10, r2. Patella I, tarsi I and II aspinose.

LEGS. Tibia and metatarsus I as shown in Fig. 132 View Figs 125–132 . Scopula entire and distal on metatarsi I and II, very narrowly divided on tarsi I and II, widely divided on tarsi III, absent on tarsi IV. Trichobothria: 2 rows of 9–11 each on tibiae; 10–12 on metatarsi; 16 on tarsi I and II; 13 on tarsi III and IV, 9–10 on cymbium. PTC I–II with 7 teeth on each margin; PTC III–IV with 7 teeth on outer and 6 teeth on inner margins.

PALP. Tibia, cymbium and palpal organ as in Figs 187 View Figs 173–187 , 220–221, 223 View Figs 215–223 . Tibia cylindrical with numerous thick ventral bristles ( Fig. 187 View Figs 173–187 ). Embolus tapering, slightly curved, embolic tip with small membranous keel ( Figs 220–221, 223 View Figs 215–223 ).

SPINNERETS. See Fig. 289 View Figs 278–290 . PMS: length 0.51, diameter 0.22. PLS: maximal diameter 0.53; length of basal, medial and apical segments 0.90, 0.65, 0.83; total length 2.38; apical segment shortly digitiform.

Female (paratype, Varzob canyon)

HABITUS. See Fig. 26. View Figs 18–26

MEASUREMENTS. TBL 22.30, CL 8.55, CW 7.32, LL 1.12, LW 1.82, SL 4.70, SW 4.11.

COLOUR. Similar to that of male, but carapace and leg more intensive reddish coloured; dark brown dorsal abdominal pattern is even better developed; ventral brownish yellow surface of abdomen with numerous dense and tiny brown spots located irregularly between last pair of book lings to spinnerets.

PROSOMA. Clypeus and eye tubercle as in Fig. 76 View Figs 63–77 . Eye diameters and interdistances: AME 0.21(0.30), ALE 0.38, PLE 0.28, PME 0.19, AME–AME 0.24(0.19), ALE–AME 0.25(0.20), ALE–PLE 0.21, PLE– PME 0.15, PME–PME 0.56. Cheliceral rastellum composed of numerous spikes located in front of fang base and on low mound. Each cheliceral furrow with 7–8 promarginal teeth and 9–13 smaller retrolateral teeth (as in Fig. 87 View Figs 78–89 ). Sternum, labium and maxillae as shown in Fig. 115 View Figs 108–116 . Sternal sigilla small, posterior pair broadly oval and located distantly from sternal margin. Maxillae with 10–11 cuspules each.

LEGS. Scopula entire and distal on metatarsi I and II, entire on palpal tarsus, very narrowly divided on tarsi I and II, elsewhere absent. Trichobothria: 2 rows of 8–9 each on tibiae; 15–18 on metatarsi; 12–17 on tarsi. Palpal claw with 4 promarginal teeth. PTC I–II with 5–6 teeth on outer and 4–5 teeth on inner margins; PTC III with 4–5 and 2–3; PTC IV with 3–4 and 1–2 teeth, respectively.

SPINATION. Palp: femur pd1; patella p1; tibia p3, v12–14; tarsus v3. Leg I: femur pd1; tibia p3, v6–7; metatarsus v5–7. Leg II: femur pd1; patella p1; tibia p3, v8–9; metatarsus v9–10. Leg III: femur r2; patella p3; tibia p2, v7; metatarsus d1, p3, r3, v9–10; tarsus p2. Leg IV: tibia v7–8; metatarsus p4, v11–12; tarsus pv7–8. All femora with 3–5 long bristles located medially, femora III and IV also with prodorsal and retrodorsal rows of bristles, with 2–3 bristles in each row; patellae I and IV, and tarsi I and II aspinose.

SPERMATHECAE. Entire and mushroom-shaped with wide tapering stalks and rounded receptacles ( Fig. 259 View Figs 245–265 ).

SPINNERETS. See Fig. 290 View Figs 278–290 . PMS: length 0.79, diameter 0.48. PLS: maximal diameter 1.02; length of basal, medial and apical segments 1.52, 0.75, 0.97; total length 3.24; apical segment triangular to shortly digitiform.

Variation

The length of the carapace varies from 6.48 to 6.97 in males and from 6.89 to 9.47 in females. The colour varies narrowly from somewhat lighter and more reddish to an even darker brown than that of the holotype. The thoracic fovea are sometimes slightly recurved. Variations in shape, arrangement and structure are shown for the eyes ( Figs 61–62 View Figs 48–62 , 76–77 View Figs 63–77 ), the cheliceral furrow ( Figs 87–89 View Figs 78–89 ), the sternum ( Figs 115–116 View Figs 108–116 ), the male tibia and the metatarsus I ( Figs 131–132 View Figs 125–132 ), the male palpal tibia and the cymbium ( Figs 186–187 View Figs 173–187 ), the palpal organ ( Figs 220–223 View Figs 215–223 ), and the spermathecae ( Figs 258–265 View Figs 245–265 ). The number of maxillary cuspules ranges from 10 to 14.

Habitat

Anemesia infuscata sp. nov. seems to be the most mesophilic congener inhabiting shrubby and forested slopes between 1000 and 2300 m ( Figs 324–329 View Figs 322–329 ).

Distribution

Tajikistan. See Fig. 373 View Fig .

Notes

Some illustrations of Anemesia infuscata sp. nov. are based on the additional paratype specimens; these include the images showing variations (see above) and photos of live spiders presented in Figs 38–41 View Figs 34–41 and 45–47 View Figs 42–47 .