Phasia rufiventris ( Macquart, 1851 ) Sun & Marshall, 2003

Sun, Xuekui & Marshall, Stephen A., 2003, Systematics of Phasia Latreille (Diptera: Tachinidae), Zootaxa 276 (1), pp. 1-320 : 181-184

3.6.14 Phasia rufiventris ( Macquart, 1851) , comb. nov.

(Figures I­53, II­8.2)

Hyalomyia rufiventris Macquart, 1851: 188 (215); ­ Crosskey 1971: 272 (type revision).

Austrophasia rufiventris: Townsend 1916: 45 ; ­ 1938: 41.

Alophora aureiventris Curran, 1927: 165 , syn. nov.

Hyalomyia costalis Malloch, 1929: 284 , syn. nov.

Hyalomyia nigrisquama Malloch, 1929: 110 , syn. nov.

Hyalomyia chrysis Malloch, 1930: 95 , syn. nov.

Hyalomyia discalis Malloch, 1930: 95 , syn. nov.

Alophora (Alophorella) aureiventris: Crosskey 1973: 110 (catalog) ­ Cantrell and Crosskey 1989: 735 (catalog) ­ Cantrell 1988: 89 (description of male and female terminalia).

Alophora (Alophorella) rufiventris: Crosskey 1973: 110 (catalog) ­ Cantrell and Crosskey 1989: 735 (catalog).

Alophora (Alophorella) costalis: Crosskey 1973: 110 (catalog) ­ Cantrell and Crosskey 1989: 735 (catalog).

Alophora (Alophorella) discalis: Crosskey 1973: 110 (catalog) ­ Cantrell and Crosskey 1989: 735 (catalog).

Alophora (Alophorella) chrysis: Crosskey 1973: 110 (catalog) ­ Cantrell and Crosskey 1989: 735 (catalog).

Alophora nigrisquama: Crosskey 1973: 110 (catalog) ­ Cantrell and Crosskey 1989: 735 (catalog).

Alophora aureiventris: Cantrell 1986: 259 (host).

DESCRIPTION

Body length: 4.5­9 mm.

MALE: HEAD. Head compressed anteriorly. Eyes separated by a distance as wide as single ocellus. Fronto­orbital plate brown to black with grey pruinosity, 1­2 rows of hairs laterally (closed to frontal setae). Frontal vitta brown, triangular. Frontal vitta at base of antennae 2.1­2.2 times as wide as fronto­orbital plate anteriorly. Ocellar setae present but fine; outer vertical setae absent or present; inner vertical setae absent. Face brown to black with grey pruinosity. Parafacial brown to black with grey pruinosity, bare, 1.1 times as wide as first flagellomere. Lower margin of face projecting, visible in profile, brown, grey pruinose. Vibrissa well differentiated; intervibrissal distance 1.1 times distance between vibrissa and eye on same side; facial ridge with bristles on lower 1/3. Gena brown, grey pruinose; hairs white; height 0.15­0.18 times eye height. Lunule black and shining; sublunular bulla indistinct. Antenna black; first flagellomere 1.1­1.2 times as long as pedicel; arista thickened on basal 0.66. Length of oral opening 2.1­2.4 times its width. Occiput flattened, white pruinose; hairs white. Palpus brown.

THORAX. Mesoscutum strongly pruinose, with 4 black longitudinal vittae, the central two vittae ended at middle of postsutural area, or merged into big black area, with fine black hairs. 0+1(2) acrostichal setae; 0(1)+1 dorsocentral setae; 2­3 postpronotal setae; presutural supra­alar seta present, strong; 1 postsutural intra­alar seta; 2 notopleural setae; 1 supra­alar seta; 2 postalar setae. Pleuron thinly grey pruinose; hairs black and white and yellow. Anepimeral setae medium size, black; 2 katepisternal setae; 6­7 yellow or black setae. Scutellum black with grey pruinosity and yellow pruinose, with two pairs of marginal setae; apical setae present and distinct; discal setae randomly arranged, not in rows. Subscutellum not very prominent. WING. Lower calypter white with light brown infuscation. Wing base without scale­like setae. Tegula brown to black. Basicosta yellow. Wing hyaline or pictured in various patterns, narrow or broad; petiole of apical cell 0.20­0.22 times as long as preceding section of R 4+5; M meeting R 4+5 almost at right angle. Halter brown. LEGS. Fore femur black; ventrally and laterally with whitish yellow hairs, dorsal surface with black hairs. Fore tibia black, without bristles, apically with 1 p and 1 d. Fore tarus black. Fore claws brown, apex black; 1.1­1.2 times as long as fifth tarsomere. Pulvilli brown. Mid femur black; hairs black, or ventrally and laterally with whitish yellow hairs, dorsal surface with black hairs. Mid tibia black, with 1 ad and 1 v, apically with 1 ad, 1 a, 1 av, 1 pv, 1 p, 1 pd, 1 v and 1 d. Hind femur black; hairs black; scale­like setae absent. Hind tibia black, with 6­8 ad and 4­5 pd.

ABDOMEN. Abdominal tergites black, or with yellow spot or area; pruinosity present, but thin; longitudinal vitta absent, or distinctly present. Relative length of abdominal tergites I+II:III:IV:V = 1.2:1:1:1.2; relative width I+II:III:IV:V = 0.4;1:1:0.9. Syntergite I+II shiny, not pruinose. Tergite III shiny. Pruinosity of tergite IV silvery, or yellow; hair spots indistinct. Pruinosity of tergite V yellow. TERMINALIA. Syncercus without notch posteriorly. Surstylus longer than cercus, strongly arched dorsally. Ejaculatory apodeme small, knob­like. Hypandrium broad, longer than phallapodeme. Epiphallus distinct. Pregonite well developed, with fine hairs dorsally. Postgonite as long as pregonite, apex spherical. Phallus broad, almost as long as hypandrium, without fine hairs dorsally. Distiphallus branched, membranous.

FEMALE: Usually smaller than male. Wing hyaline. Eyes touching. Anterior spinelike setae of hind tibia absent. Abdominal tergites black or with yellow transverse bands. Sternite VII (sheath) longer than sternite VI, slender, pointed apically, slightly bent, apex directed dorsally, sternite VII with linear wrinkles ventrally. Ovipositor bent upward.

TYPE MATERIAL

Hyalomyia rufiventris Macquart, 1851 . Holotype ♂, [ AUSTRALIA], Tasmania, No. 2308, "3 47" (MNHN, Paris, examined).

Alophora aureiventris Curran, 1927 . Holotype ♂, [ AUSTRALIA], Queensland, Biloela, Apr. 1927, ex: Dysdercu sidae Montr. , with pupa, E. Ballard (BMNH, examined); allotype ♀, AUSTRALIA, Queensland, Gatton, Oct. 1926, ex: Dysdercus sidae Montr. , with pupa, E. Ballard (BMNH, examined); paratypes, 2♂ 1♀, AUSTRALIA, Queensland, Brisbane, 10.iv.1926, ex: Dysdercus sidae, E. Ballard (QDPC, AMNH, examined).

Hyalomyia costalis Malloch, 1929 . Holotype ♂, AUSTRALIA, Woodford, 15.xi.1925, Mackerras (ANIC, examined).

Hyalomyia nigrisquama Malloch, 1929 . Holotype ♂, [ AUSTRALIA], N. Sydney, French Forest, 14.xii.1923, T.G. Campbell (AM, examined).

Hyalomyia chrysis Malloch, 1930 . Holotype ♂, [ AUSTRALIA], Western Australia, Narrogin, 30.viii.1926, E.W. Ferguson (ANIC, examined).

Hyalomyia discalis Malloch, 1930 . Holotype ♂, [ AUSTRALIA], Western Australia, Geraldton, 5.ix.1926, E.W. Ferguson (ANIC, examined).

OTHER MATERIAL EXAMINED

AUSTRALIA: Australian Capital Territory: Canberra, (1♂, ANIC); Canberra, 17.ii.1948, S.J. Paramonov (1♂, ANIC). New South Wales: Bilpin nr. Kurrajong, 23.v.1979, N.W. Rodd (1♂, AMSA); Blue Mountains, Bilpin, 2.ix.1977, 26.ix.1978, N.W. Rodd (6♂, AMSA); Blue Mountains, Mt. Tomah, 26.x.1978, 12.iii.1981, N.W. Rodd (2♂, AMSA); Blue Mountains, Mt. York, 20.i.1982, N.W. Rodd (1♂, AMSA); Kanangra, Boyd National Park, Budthingaroo Cr., 25.xii.1977, G. Daniels (1♂, AMSA); Cranmore Park [?], 12. xi.1933, Fuller (1♂, ANIC); Deepwater (NE), Cox's Rd. Forest lands s.f. 17.xii.1985, G. Williams (1♂, AMSA); Galston Gorge, 27.viii.1959, M. Nikitin, B.M. 1959­616 (1♂, BMNH); Jnverell [?], 29.viii.1959, J. Crothers (1♂, NSWA); Kinchega National Park, 11­ 13.ix.1980, N.W. Rodd (1♂, AMSA); 7.5 mi E Braidwood, Mongarlowe R., 1.xii.1972, G.B. Fairchild (2♂, USNM); Mt. Donna Buang [?]. v. and 3­2­1951 (1♂, MVNH); Sydney, (5♀, NSWA); Waratah track, Gibraltar Range National Park, 10­12.xi.1984, D.K. Yeates (1♂, UQIC); Yass, 16­17.x.1935, 14.xi.1939, K. English (1♂ 2♀, AMSA, NSWA). Queensland: Biloela, 3.v.1927, ex: Dysdercus sidae , (1♂, BMNH); Gladstone (SW 65km), Kroombit Tops, 24.ii.1982, D. Yeates (1♂, UQIC); Gatton, 3.viii and 19.x.1927, M.S. Evans (1♂ 1♀, QDPC, BMNH); Laidley, 2.ii.1928, ex: Dysdercus sidae, M.G. Evans (3♂ 1♀, BMNH, with pupae); Laidley, 18.x.1927, ex: Dysdercus sidae, M.S. Evans (1♂ 2♀, QDPC, BMNH); Sydney, Manly, 6.12.1923 (1♂, BMNH); Sunny Bank, 18.ix.1927 (1♀, UQIC); Tamborine Mts. (SE), 18­25.v.1935 (3♂ 1♀, BMNH); Yarraman (SE, 7 km W), 17.iv.1976, host: Oncopeltus sordidus [Dallas], A. Slater (1♂ 1♀, QDPC). South Australia: Adelaide, v.1962, C. Watts (1♂, CNCI); Edillilie (10 mi N), 7.xii.1968, N. McFarland (4♂, SAMA); Rev. [?], A.P. Burgess (1♂, SAMA). Tasmania: Mangalaze [?], 1908, A. White, B.M. 1917­104 (1♂, BMNH). Victoria: Princetown, Ocean Rd., 27.xi.1977, J.F. Donaldson (1♀, QDPC); Warburton, 9.12.18 (1♀, MVNH). Western Australia: Beverley (1♂, SAMA); Capel, 7.1.1957, A. Snell (3♀, MVNH); Dongara, 26.x­3.x.1935, R.E. Turner, B.M. 1935­240 (1♀, BMNH); Dumbleyung, 27­30.x.1963, H. Udell (3♂ 1♀, WAMP); Slonevilla (17km N), Berry Res. [?], 11.xi.1992, T.F. Houston and R.P. McMillan (1♂, WAMP).

HOSTS

Oncopeltus sordidus ( Hemiptera , Lygaeidae ): ­ Australia ( Cantrell 1986: 259).

Dysdercus sidae Montrouzier ( Hemiptera , Pyrrhocoridae ): ­ Australia ( Crosskey 1973: 179).