Euthria frausseni, Harzhauser & Landau, 2024

Harzhauser, Mathias & Landau, Bernard M., 2024, The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5427 (1), pp. 1-110 : 75-76

Euthria frausseni nov. sp.

Figs 34F , 40A–C

? Euthria adunca ( Bronn, 1831) — Kovács & Vicián 2023: 253, fig. 13A [non Euthria adunca ( Bronn, 1831) ].

Type material. Holotype: NHMW 1865 /001/0207, SL: 29.1 mm, MD: 12.2 mm, Lăpugiu de Sus ( Romania), Figs 40A 1 –A 2 . Paratypes: NHMW 1870 /0033/0097, SL: 29.0 mm , MD: 13.1 mm, Lăpugiu de Sus ( Romania), Figs 40B 1 – B 2 . NHMW 1854 /0035/0210, SL: 25.9 mm , MD: 11.7 mm, Lăpugiu de Sus ( Romania), Figs 40C 1 –C 2 . NHMW 2023 /0338/0012, SL: 24.5 mm , MD: 10.9 mm, Lăpugiu de Sus ( Romania), Figs 34F .

Additional material. 15 spec., NHMW 1866 /0045/0299, Lăpugiu de Sus ( Romania) .

Type locality. Lăpugiu de Sus ( Romania)

Type stratum. Silt and clay of the Dej Formation.

Etymology. In honor of Koen Fraussen ( Belgium), in recognition of his contributions on Buccinoidea.

Age. Middle Miocene, early/middle Badenian (Langhian).

Diagnosis. Medium-sized, moderately slender fusiform shell with low, dome-shaped, paucispiral protoconch and moderately high spire of strongly convex whorls, sculpture changing from prominent axial and spiral sculpture on early teleoconch whorls to relatively smooth on third to fourth teleoconch whorl, base strongly constricted, siphonal canal moderately long, narrow, moderately recurved.

Description. Medium-sized, moderately slender fusiform shell of up to seven teleoconch whorls; apical angle ~45°. Protoconch paucispiral, low dome-shaped of 1.25 whorls; diameter: 800 μm, height: 600 μm. Early teleoconch whorls convex with periphery at abapical suture, bearing broad axial ribs, separated by narrower interspaces, overrun by four primary spiral cords with delicate secondaries intercalated. Suture shallowly undulating around ribs. Axial ribs fading on third to fourth teleoconch whorl. Later teleoconch whorls moderately convex, with periphery below mid-whorl, separated by moderately incised suture, bearing faint spiral threads. Last whorl attaining ~65% of total height, evenly convex, smooth, base strongly constricted, bearing weak spiral cords, fasciole indistinct. Aperture moderately wide, pyriform. Columella deeply excavated, angled at transition to siphonal canal; transition often accentuated by small denticle. Columellar callus indistinct, poorly delimited from base. Columellar callus smooth or with up to three small denticles in abapical part. Anal canal broad, accentuated by small distinct parietal denticle. No anal denticle. Outer lip thin at peristome, with prominent lirae placed some distance behind peristome, passing into about 12 delicate lirae extending deep within aperture lirae. Siphonal canal moderately long, narrow, deflected to the left, moderately recurved, shallowly notched.

Discussion. This species is closely similar to Euthria walaszczyki nov. sp. and the adult shells of the two are strikingly similar. However, they differ by the constantly smaller size of Euthria frausseni nov. sp. and especially by their protoconchs and early teleoconchs. The protoconch is paucispiral low dome-shaped in E. frausseni as opposed to multispiral high conical in E. walaszczyki and its early teleoconch whorls lack the shoulder of E. walaszczyki . This species had been misidentified as Euthria intermedia ( Michelotti, 1847) in the collection of the NHMW, but differ from that Early Miocene Italian species by the smooth, convex late spire whorls, the longer siphonal canal and the lack of prominent, wide-spaced spiral cords on the base.

Paleoenvironment. Unknown, probably middle to outer neritic environments.

Distribution in Central Paratethys. Badenian (Middle Miocene): North Alpine Foreland Basin: Grund ( Austria) (Hoernes & Auinger 1890); Pannonian Basin: Letkés ( Hungary) ( Kovács & Vicián 2023); Făget Basin: Lăpugiu de Sus ( Romania) (Hoernes & Auinger 1890).

Figures

FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width.

FIGURE 2. Colubraria subobscura (Hoernes & Auinger, 1884). A1–A3. NHMW 1854/0035/0170, paralectotype, Lăpugiu de Sus (Romania). B1–B2. NHMW 1859/0037/0018a, paralectotype, Lăpugiu de Sus (Romania). C1–C3. NHMW 1865/0001/0183, paralectotype, Lăpugiu de Sus (Romania). D1–D2. NHMW 1859/0037/0018b, paralectotype, Lăpugiu de Sus (Romania). E1–E3. NHMW 1874/0025/0021, lectotype, Lăpugiu de Sus (Romania). F1–F2. NHMW 2023/0338/0001, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 34.A. Euthria brunettii nov.sp., NHMW2023/0338/0007,paratype, Lăpugiu de Sus(Romania).B.Euthria walaszczyki nov. sp., NHMW 1868/0001/0223, Grund (Austria). C. Euthria depressospira Bandat, 1943, NHMW 2023/0338/0008, Lăpugiu de Sus (Romania). D. Euthria diluvii (Eichwald, 1830) NHMW 1860/0001/0224b, Boršov (= Porstendorf) (Czech Republic). E. Euthria fuscocingulata (Hoernes & Auinger, 1890). NHMW 2023/0338/0011, paralectotype, Lăpugiu de Sus (Romania). F. Euthria frausseni nov. sp., NHMW 2023/0338/0012, paratype, Lăpugiu de Sus (Romania). G. Euthria odiosa nov. sp. NHMW 2023/0348/0001, paratype, Steinebrunn (Austria). H. Euthria puschii (Andrzejowski, 1830), NHMW 2023/0350/0002, Bad Vöslau (Austria). I. Euthria stuetzii (Naumann, 1852), NHMW 1846/0037/0254, Enzesfeld or Gainfarn (Austria). J. Euthria subnodosa (Hoernes & Auinger, 1890), NHMW 1846/0037/0274, Steinebrunn and Gainfarn (Austria). K. Euthria yesimae nov. sp., RGM 794 572, paratype, Pınarlar Yaylası, Akpınar (Turkey).

FIGURE 40. Euthria frausseni nov. sp. A1–A2. NHMW 1865/001/0207, holotype, Lăpugiu de Sus (Romania). B1–B2. NHMW 1870/0033/0097, paratype, Lăpugiu de Sus (Romania). C1–C2. NHMW 1854/0035/0210, paratype, Lăpugiu de Sus (Romania).