Coilodera formosana Moser, 1910

Qiu, Jian-Yue, Xu, Hao & Chen, Li, 2017, Taxonomic review of the Oriental flower beetle Coilodera penicillata species complex (Coleoptera: Scarabaeidae: Cetoniinae), Zootaxa 4350 (3), pp. 511-537 : 524-525

publication ID

https://doi.org/ 10.11646/zootaxa.4350.3.5

publication LSID

lsid:zoobank.org:pub:7B3B84E2-4521-4591-8F53-71F56018A403

DOI

https://doi.org/10.5281/zenodo.6040981

persistent identifier

https://treatment.plazi.org/id/03CD6B0A-FFD9-FC44-2E81-FB7FF73EF8C4

treatment provided by

Plazi

scientific name

Coilodera formosana Moser, 1910
status

 

Coilodera formosana Moser, 1910 , revised status

( Figs. 25–28 View FIGURES 21 – 33 , 45–46 View FIGURES 34 – 48 , 57, 65, 71 View FIGURES 49 – 72 , 81–82, 85 View FIGURES 73 – 85 )

Coelodera nigroscutellaris formosana Moser, 1910: 297 View in CoL (type locality: Formosa [Tainan, Taiwan]); Schenkling 1921: 133; Miwa 1931: 305; Miwa & Chûjô 1939: 83 (synonym of C. penicillata Hope ).

Coelodera penicillata formosana Moser View in CoL : Mikšić 1972: 703, fig. 20; Mikšić 1976: 271, fig. 33a; Miyake et al. 1991: 39; Yu et al. 1998: 101, 180, plate 6, figs. 1a ♂, 1b ♀.

Coilodera penicillata formosana Moser : Krajčík 1998: 88; Sakai & Nagai 1998: 358 (Nantou Hsien, Taiwan), plate. 136, figs. 1625-4 ♂, 1625-5 ♀; Smetana 2006: 306; Krajčík 2012: 76; Bezděk 2016: 401.

Coelodera penicillata (nec Hope): Kobayashi 1980: 20 (misapplication of the name).

Macronota hopponus Niijima & Kinoshita, 1923: 232 View in CoL (type locality: Hoppo, Formosa [=Peipu, Hsinchu Hsien, Taiwan), plate. I, fig. 12. Synonymized with C. penicillata by Miwa & Chûjô 1939: 83.

Diagnosis. See remarks of C. penicillata .

Variation. Body length 23.6–24.9 mm; width 10.7–11.4 mm. Most individuals with orange-red tomentum ( Fig. 25, 28 View FIGURES 21 – 33 ) and wine-red metallic reflections, while a small number with yellow tomentum and green metallic reflections ( Figs. 26–27 View FIGURES 21 – 33 ). A few males with setae in the abdominal groove ( Fig. 28 View FIGURES 21 – 33 ).

Type material. Syntype of Coilodera formosana labeled: Formosa, Tainan [handwriting on pale yellow label] / formosana Mos. Type [handwriting of Moser on pale yellow label] / Type [red label] / subsp. formosana Mos. [handwriting of Moser on pale yellow label] (female, MFNB, Figs. 45–46 View FIGURES 34 – 48 ).

Comments on type material. The sex, number, and collecting site of the type series of Coilodera formosana were not mentioned in the original publication ( Moser 1910). Therefore, it was difficult to determine whether other specimens from Formosa in MFNB were included in the type series, and only the single female with a handwritten identification label of Moser here regard as syntype.

The type specimens (one male and three female) of Macronota hopponus Niijima & Mastumura, 1923 are housed in EIHU (Dr. Masahiro Ohara, personal communication, September 2014) and unavailable for study at present.

Other material examined (11♂♂, 24♀♀). Taiwan: 1♂ ( BMNH), Horisha [ Puli , Nantou Hsien], Col. T. Shiraki ; 1♀ ( SYSM), Formosa Hori ( Puli Polisia ) Taichung District. 800 m, 22.VIII.1947, L. Gressitt ; 1♀ ( SYSM), Formosa Hori ( Puli Polisia ) Taichung District. 800 m, 23.VIII.1947, L. Gressitt ; 1♂ ( QCCC), 30.IV.2007, Mount Guandaoshan , Puli, Nantou Hsien, Chang-Chin Chen ; 2♀♀ ( MFNB), Puli , V.1961, C Formosa ; 2♂♂ ( MFNB), Puli , 10.VI.1964, C Formosa ; 2♂♂ ( MFNB), Hori [ Puli , Nantou Hsien], 16.VIII.1935, Formosa ; 1♀ ( MNHN), Hokuzanko [Peishankeng], Hori , Formosa, V.1938, U. Shiga ; 1♂ ( MHNG), Puli , Formosa // V.1964 ; 1♂, 1♀ ( QCCC), 7.X.2012, Xiama, Haiduan, Taitung Hsien, Yu-Tang Wang ; 1♀ ( QCCC), 23.V.2012, Zhilexi, Taichung Hsien, Fu-Sheng Huang ; 1♀ ( QCCC), km 74 of Alishan Highway , Chiayi Hsien, 15.VII.2014, Chao-SenYang ; 1♀ ( QCCC), 16.VII–5.VIII.2004, Shih Lin , Taoyuan Hsien, ~ 1,180 m, flight intercept trap, Chun-Lin Li ; 1♀ ( QCCC), 1–13.VII.2010, Shih Lin , Taoyuan Hsien, ~ 1,180 m, flight intercept trap, Chun-Lin Li ; 1♂ ( BMNH, Bourgoin Coll.), Kosempo [ Chiasien , Kaohsiung Hsien], Formosa, H. Sauter, 1911 // 7.VII ; 1♂ ( SDEI), Kosempo ( Formosa), H. Sauter, V.1912 ; 1♀ ( SDEI), Kosempo ( Formosa), H. Sauter, 1912 ; 1♀ ( SDEI), Kosempo, Formosa, H. Sauter, 1911 ; 1♀ ( BMNH, Bourgoin Coll.), Hoozan [ Fengshan , Kaohsiung Hsien], Formosa, H. Sauter, 1911 // 7.IX ; 1♀ ( MFNB), Formosa, Hoozan , IX.10, H. Sauter S. G. ; 1♀ ( MFNB), Taiheizan [Taipingshan, Ilan Hsien] ( Formosa), Makurei , 18.VII.1940, S. Miyamoto ; 1♀ ( SDEI), Chip Chip [Chichi, Nantou Hsien], Formosa, Sauter , X.1908 ; 1♀ ( SDEI), Bukai [ Wuchieh , Nantou Hsien], Formosa, 13.VI.1934 ; 1♀ ( NHMB), Formosa, Coll. Dr. Itzinger ; 1♀ ( NHMB), Formosa // Sammlung Schürhoff ; 1♀ ( NHMB), Tamari [ Taimali ], Taito [Taitung Hsien], Formosa // Sammlung Schürhoff ; 1♀ ( MFNB), Tamari, Taito, Formosa ; 1♀ ( MFNB), Formosa, Sauter ; 1♂, 1♀ ( MNHN), Hoppo , Formosa, V.1960 ; 1♀ ( BMNH), 16.XI.2008, Taitung County, 14 km W of Chihshang , 23°09′N, 121°04′E, 900 m, L. Dembicky, BMNH 2008-85 GoogleMaps ; 1♀ ( BMNH, Bourgoin Coll.), Suiharyo , Formosa, H. Sauter, 1911 // 7.XII.

Distribution. This species is endemic to the island of Taiwan.

Natural history. A newly emerged adult of this species was recorded in the wood-feeding cockroach galleries of Salganea raggei (Blattaria: Blaberidae : Panesthiinae ) ( Kon et al. 2003).

Remarks. The larvae morphology of Coilodera formosana and seven other species of Taenioderini were presented in detail in Vendl et al. (2014), which is the first study on the immature stages of this tribe.

MFNB

Museo Friulano di Storia Naturale

MNHN

Museum National d'Histoire Naturelle

MHNG

Museum d'Histoire Naturelle

NHMB

Naturhistorisches Museum, Basel

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Cetoniidae

Genus

Coilodera

Loc

Coilodera formosana Moser, 1910

Qiu, Jian-Yue, Xu, Hao & Chen, Li 2017
2017
Loc

Coilodera penicillata formosana

Bezdek 2016: 401
Krajcik 2012: 76
Krajcik 1998: 88
Sakai 1998: 358
1998
Loc

Coelodera penicillata

Kobayashi 1980: 20
1980
Loc

Coelodera penicillata formosana

Yu 1998: 101
Miyake 1991: 39
Miksic 1976: 271
Miksic 1972: 703
1972
Loc

Macronota hopponus

Miwa 1939: 83
Niijima 1923: 232
1923
Loc

Coelodera nigroscutellaris formosana

Miwa 1939: 83
Miwa 1931: 305
Schenkling 1921: 133
Moser 1910: 297
1910
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