Ceratophyus, Fischer von Waldheim, 1824

Král, David, Jákl, Stanislav, Lu, Yuanyuan & Bai, Ming, 2020, The genus Ceratophyus Fischer von Waldheim, 1824 (Coleoptera: Geotrupidae) in China, along with description of a new species, Zootaxa 4747 (2), pp. 339-349 : 345-346

Key to males of eastern Palearctic Ceratophyus species

Species distribution after Hillert & Král (2013), Nikolajev et al. (2016) and this paper.

1 Apical tooth of protibia simple; aedeagus as in Figs. 7–9 ; body length 18.0– 23.5 mm; China ( Hebei, Heilongjiang, Inner Mongolia), Mongolia ( Dornod, Hentiy, Sükhbaatar), Russia ( Buryatiya, Zabaykalskiy Kray)..................................................................................................... Ceratophyus dauricus ( Jekel, 1866)

- Apical tooth of protibia bifid. …......................................................................... 2

2 Elytral striae finely impressed, becoming finer to absent posteriorly, elytral intervals flat; aedeagus as in Figs. 10–12 ; body length 19.5–21.0 mm; China ( Sichuan). ….................................... Ceratophyus kabaki Nikolajev, 2007

- Elytral striae distinctly impressed, visible to apex, elytral intervals convex ( Fig. 1 ).................................. 3

3 Base of pronotal horn and areas laterally of it rugose; aedeagus as in Figs. 16–18 ; body length 18.0– 27.5 mm; China ( Beijing, Gansu, Hebei, Inner Mongolia, Ningxia, Shaanxi, Shanxi, Qinghai and Xinjiang), Mongolia ( Dornogovi)......................................................................................... Ceratophyus sinicus Zunino, 1973

- Base of pronotal horn and areas laterally of it simply punctate.................................................. 4

4 Setation of lateral margin of head, pronotum and ventral surface reddish brown; areas laterally of pronotal horn smooth (without punctures), elytral stria 5 usually reaching elytral base; aedeagus as in Figs. 13–15 ; body length 18.0– 27.5 mm; China ( Xinjiang), Kazakhstan, Kyrgyzstan............................... Ceratophyus mesasiaticus Medvedev & Nikolajev, 1974

- Setation of lateral margin of head, pronotum and ventral surface blackish ( Figs 1–3 ); whole surface of pronotal disc distinctly, densely punctate ( Fig. 1 ), elytral stria 5 usually not reaching base ( Fig. 1 ); aedeagus as in Figs. 4–6 ; body length 18.5–22.0 mm; China ( Gansu, Qinghai)........................................... Ceratophyus amdoensis Král & Jákl , new species

Figures

FIGURES 4–12. Aedeagus of Ceratophyus Fischer von Waldheim, 1824 species. 4–6, Ceratophyus amdoensis Král & Jákl, new species, holotype; 7–9, C. dauricus (Jekel, 1866) (“Transbaikalia”, NMPC); 10–12, C. kabaki Nikolajev, 2007 (Sichuan, Yading, NMPC). 4, 7, 10, dorsal view; 5, 8, 11, left lateral view; 6, 9, 12, tips, ventral view. Not to scale.

FIGURES 1–3. Ceratophyus amdoensis Král & Jákl, new species, habitus of holotype (♂). 1, dorsal view; 2, ventral view; 3, left lateral view. Not to scale.

FIGURES 13–18. Aedeagus of Ceratophyus Fischer von Waldheim, 1824 species. 13–15, C. mesasiaticus Medvedev & Nikolajev, 1974 (Kazakhstan, Karatal, NMPC); 16–18, C. sinicus Zunino, 1973 (Qinghai, Haibu env., NMPC). 13, 16, dorsal view; 14, 17, left lateral view; 15, 18, tips, ventral view. Not to scale.