Physotarsus Townes, 1966

Zhaurova, Kira & Wharton, Robert, 2009, A revision of Physotarsus Townes (Hymenoptera: Ichneumonidae: Ctenopelmatinae), with description of 18 new species, Zootaxa 2207, pp. 1-52 : 9-10

Key to species of Physotarsus Townes, 1966

1 Fore wing weakly ( Fig. 9 ) to strongly ( Fig. 44 ) yellowish or brown to black at base, with ( Figs 9 , 32 ) or without ( Figs 46 , 61 ) infumate areas apically .......................................................................................................................... 2

- Fore wing hyaline at base (as in Figs. 13, 15 ) ............................................................................................................. 8

2(1) Fore wing evenly colored, entirely infumate/fuscous ( Fig. 46 ) or pale yellowish-brown ......................................... 3

- Fore wing not evenly colored (as in Figs 44 , 61 ) ....................................................................................................... 4

3(2) Fore wing pale yellowish brown .......................................................................................... P. albus Zhaurova , n. sp.

- Fore wing dark, entirely fuscous .................................................................................. P. m e l i p e n n i s Zhaurova, n. sp.

4(2) Basal part of fore wing dark brown to black, apical part variable ( Figs 44 , 61 ) ........................................................ 5

- Basal part of fore wing yellowish, apex infumate (as in Figs 9 , 32 ) .......................................................................... 6

5(4) Mesosoma dorsally, excluding pronotum, orange; hind femur and tibia orange ..... P. maculipennis ( Cresson, 1874)

- Mesosoma dorsally, excluding pronotum, dark brown to black; hind femur and tibia black ....................................... ....................................................................................................................................... P. varicornis ( Cameron, 1886)

6(4) Mesosoma with yellow and black markings, face quite densely punctate medially ( Fig. 33 ) .................................... ....................................................................................................................................... P. flavipennis Zhaurova, n. sp.

- Mesosoma entirely yellowish to light orange, face very lightly evenly punctate ( Fig. 28 ) ........................................ 7

7(6) Frons and first tarsomere of hind leg orange ............................................................................ P. adriani Gauld, 1997

- Frons and first tarsomere of hind leg black ............................................................................ P. castilloi Gauld, 1997

8(1) Epicnemial carina strongly turning towards anterior margin of mesopleuron, reaching it or nearly so .................... 9

- Epicnemial carina largely parallels anterior margin of mesopleuron ....................................................................... 14

9(8) T1 with apical third white, otherwise largely dark brown; ocellar field entirely black .. P. oculatus Zhaurova , n. sp.

- T1 either uniformly colored or dark apically, at most with narrow white apical margin; ocellar field variable, entirely black in some species, not in others .......................................................................................................................... 10

10(9) T1 entirely yellowish, orange, or light brown, color even ..................................................................................... 11

- T1 partly to entirely dark brown .............................................................................................................................. 12

11(10) T1 2.3X as long as broad ....................................................................................................... P. eli ethi Gauld, 1997

- T1 1.2X as long as broad ................................................................................................... P. j a m e s i Zhaurova, n. sp.

12(10) Propodeum brown, at least on anterior half ..................................................................... P. niveus Zhaurova , n. sp.

- Propodeum entirely orange ....................................................................................................................................... 13

13(12)Clypeus yellowish orange; hind femur dark brown ......................................... P. leucohypopygus Zhaurova , n. sp.

- Clypeus white; hind femur yellowish orange ................................................................. P. glabellus Zhaurova , n. sp.

14(8) Ventral clypeal margin with a prominent angulate central lobe ( Fig. 7 ) ................................................................ 15

- Ventral clypeal margin not as above ......................................................................................................................... 16

15(14) Body entirely yellowish orange with no brown or black markings ..................................... P. bonillai Gauld, 1997

- Mesoscutum and face with dark brown markings; anterior half of the propodeum brownish to black ....................... ............................................................................................................................................ P. claviger Zhaurova, n. sp.

16(14) Mesosoma entirely yellow to orange .................................................................................................................... 17

- Color of mesosoma variable, but at least partly black .............................................................................................. 19

17(16) Petiole dark brown ...................................................................................................... P. glabellus Zhaurova , n. sp.

- Petiole either pale yellow or dark orange ................................................................................................................. 18

18 (17) Petiole pale yellow; antenna abruptly changing from pale to dark subapically ............. P. luteus Zhaurova , n. sp.

- Petiole dark orange; antenna mostly dark throughout, without abrupt color change ......... P. j a m e s i Zhaurova, n. sp.

19(16) Face quite strongly protruding medially ( Figs 49 , 55 ) ......................................................................................... 20

- Face not to very weakly protruding medially ( Fig. 24 ) ............................................................................................ 23

20(19) Mesoscutum largely impunctate posteriorly, very sparsely punctate anteriorly ............ P. gineus Zhaurova , n. sp.

- Mesoscutum densely punctate anteriorly and posteriorly ......................................................................................... 21

21(20) Mesoscutum black with anterolateral margin pale yellowish; fore and mid femora partially black ........................ ............................................................................................................................................. P. t o n i c u s Zhaurova, n. sp.

- Mesoscutum black with anterolateral margin and most or all of notaular lines pale to dark yellow; fore and mid fem- ora yellowish ............................................................................................................................................................. 22

22(21) Mesopleuron entirely black; face yellowish laterally, black medially ......................... P. foveatus Zhaurova , n. sp.

- Mesopleuron black with yellow markings; face entirely yellowish ............................ P. melotarsus Zhaurova , n. sp.

23(19) Mesopleuron and mesoscutum entirely black ........................................................ P. montezuma ( Cameron, 1886)

- Mesopleuron usually and mesoscutum always with at least some yellow ............................................................... 24

24(23) Tarsomeres 3 and 4 of hind leg bright yellow, much lighter than tarsomeres 1 and 5. Male subgenital plate slightly emarginate ( Fig. 22 ) ......................................................................................... P. concavus Zhaurova , n. sp.

- Tarsomeres of hind leg uniformly orange or brown or grading from orange to brown. Male subgenital plate deeply emarginate ( Figs 25 , 31 , 59 ) ..................................................................................................................................... 25

25(24) Tarsomeres of hind leg distinctly infumate, at least dorsally. Antenna mostly dark brown over apical half, often with somewhat indistinct whitish subapical ring ....................................................... P. emarginatus Zhaurova , n. sp.

- Taromeres of hind leg orange. Antenna mostly orange to yellow orange over apical half, with apical flagellomeres contrastingly darker..................................................................................................................................................... 26

26(25) Mesoscutum with a narrow anterolateral yellow margin with yellow extending posteriorly along notaular line ( Fig 26 ); male subgenital plate deeply emarginate, cordate ( Fig. 25 ) ..................................... P. cordatus Zhaurova , n. sp.

- Mesoscutum with anterolateral margin usually black, never with yellow on notaular line ( Fig 60 ); male subgenital plate deeply emarginate, inner margin not protruding ( Fig. 59 ) ..................................... P. truncatus Zhaurova , n. sp.

Figures

FIGURES 9 – 12. Habitus and face. 9 – 10, Physotarsus adriani n. sp.; 9, habitus; 10, face. 11 – 12, P. albus n. sp.; 11, habitus; 12, face.

FIGURES 41 – 44. Habitus and face. 41, Physotarsus leucohypopygus, n. sp., face. 42 – 43, P. lu t eu s n. sp.; 42, habitus; 43, face. 44, P. maculipennis, habitus.

FIGURES 29 – 32. Habitus, face, and male genitalia. 29 – 31, Physotarsus emarginatus n. sp.; 29, habitus; 30, face; 31, male genitalia, ventral view. 32, P. flaivpennis n. sp., habitus.

FIGURES 45 – 48. Habitus and face. 45, Physotarsus maculipennis, face. 46 – 47, P. melipennis n. sp.; 46, habitus; 47, face. 48, P. mel ota rs us n. sp., habitus.

FIGURES 61 – 64. Habitus and face. 61 – 62, Physotarsus varicornis; 61, habitus; 62, face. 63 – 64, P. j a m e s i n. sp.; 63, habitus; 64, face.

FIGURES 13 – 16. Habitus and face. 13 – 14, Physotarsus bonillai; 13, habitus; 14, face. 15 – 16, P. castilloi; 15, habitus; 16, face.

FIGURES 33 – 36. Habitus and face. 33, Physotarsus flavipennis n. sp., face. 34 – 35, P. foveatus n. sp.; 34, habitus; 35, face. 36, P. gineus n. sp., habitus.

FIGURES 25 – 28. Physotarsus spp. 25 – 26, P. cordatus n. sp.; 25, male genitalia, ventral view; 26, head and mesonotum, dorsal view. 27 – 28, P. eliethi; 27, habitus; 28, face.

FIGURES 5 – 8. ESEM, clypeus. 5, Physotarsus concavus n. sp., 6, P. luteus n. sp., 7, P. claviger n. sp., 8, Scolobates auriculatus.

FIGURES 49 – 52. Habitus and face. 49, Physotarsus melotarsus n. sp., face. 50 – 51, P. niveus n. sp.; 50, habitus; 51, face. 52, P. oculatus n. sp., habitus.

FIGURES 53 – 56. Habitus and face. 53, Physotarsus oculatus n. sp., face. 54 – 56, P. to nicu s n. sp.; 54, habitus; 55, face Ψ; 56, face ♂.

FIGURES 21 – 24. Habitus, face, and male genitalia. 21 – 22, Physotarsus concavus n. sp.; 21, face; 22, male genitalia, ventral view. 23 – 24, P. cordatus n. sp.; 23, habitus; 24, face.

FIGURES 57 – 60. Physotarsus truncatus n. sp. 57, habitus; 58, face; 59, male genitalia, ventral view; 60, head and mesonotum, Ψ, dorsal view.