Phanaeus zoque Moctezuma & Halffter, 2017

Moctezuma, Victor & Halffter, Gonzalo, 2021, Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species, European Journal of Taxonomy 747, pp. 1-71 : 50-51

Phanaeus zoque Moctezuma & Halffter, 2017

Figs 1Q , 15 , 18R , 19R

Phanaeus zoque Moctezuma & Halffter, 2017: 47 , 48, 52, 55–56, figs 1–15, 23.

Phanaeus zoque – Moctezuma et al. 2017: 113–114, 126, 128, 131–132, fig. 19; 2019: 252–253, fig. 5. — Kohlmann et al. 2018: 83, 88.

Type material

Holotype

MEXICO • ♂, Moctezuma & Halffter 2017: 48 (revised); Oaxaca, Santa María Chimalapa, Chocomanatlán ; IEXA.

Type locality

Mexico, Oaxaca, Santa María Chimalapa, Chocomanatlán (José López Portillo).

Distribution

Temperate mountains of Los Chimalapas, eastern Oaxaca and western Chiapas ( Fig. 15 ).

Remarks

Mean length 15.8 mm (13.2–18.4 mm). To the original description we add the elytral striae impressed basally as distinct fossae; right and left lobes of endophallite copulatrix similar in size; right lobe obtusely triangular in shape, distinctly rounded superiorly; left lobe strongly developed, obtusely lobed; central ridge less developed than central column ( Fig. 1Q ). For the female, the head showing a cephalic trituberculate carina; with conical, similar in size tubercles; middle tubercle slightly more frontally projected than lateral tubercles; frons with almost effaced to effaced punctures; pronotal sculpture completely smooth, with almost effaced to effaced punctures; pronotal process trituberculate, with posterior concavity; with dentiform middle tubercle, strongly more developed and projected frontally than lateral tubercles; carinate or rounded lateral tubercles; posterior pronotal midline almost completely effaced.

After having described P. zoque ( Moctezuma & Halffter 2017) , the following paratypes were deposited by us in additional collections: CEMT: 5 ♂♂, 5 ♀♀; CDC: 5 ♂♂, 5 ♀♀; CMNC: 1 ♂, 1 ♀; CPFA: 6 ♂♂, 6 ♀♀; UVGC: 1 ♂, 1 ♀. These paratypes were previously housed at VMC (see the full list of paratypes in Moctezuma & Halffter 2017).

Figures

Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA).

Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta.

Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017.

Fig. 19. Elytral integument of species within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez- Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta). H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002.K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017.