Belisana daxiangi Li, Zheng & Yao, 2023

Zhao, Fangyu, Yang, Lan, Li, Shuqiang, Zheng, Guo & Yao, Zhiyuan, 2023, A further study on the Belisana spiders (Araneae: Pholcidae) from Xishuangbanna, Yunnan, China, Zootaxa 5351 (5), pp. 543-558 : 549

Belisana daxiangi Li, Zheng & Yao sp. nov.

Figs 4–5 , 9C–D

Type material. Holotype: Male (SYNU-Ar00310), near Bubeng Village (21°36.827’N, 101°34.847’E, elevation 690 m), Mengla County, Xishuangbanna , Yunnan, China, 12 August 2012, Guo Zheng, Xue Li, and Wenyue Zhu leg.

Etymology. The specific name is a patronym in honor of the late Chinese arachnologist Daxiang Song; noun (name) in the genitive case.

Diagnosis. The new species resembles B. fengzheni sp. nov. ( Figs 6– 7 , 9E–F , 10C–D ) having similar bulbal apophyses ( Fig. 5A ) and male chelicerae ( Fig. 5B ), but it can be distinguished by prolatero-distal membranous lamella of procursus with weakly sclerotized lamella (arrow 1 in Figs 4C , 9C ; vs. absent in B. fengzheni sp. nov.), by procursus with two spine-shaped dorso-distal apophyses (arrows 2, 3 in Figs 4C , 9C ; vs. only one apophysis in B. fengzheni sp. nov.), and by procursus with ventral membranous lamella (arrow 4 in Figs 4C , 9C ; vs. absent in B. fengzheni sp. nov.).

Description. Male (holotype, SYNU-Ar00310): Total length 1.58 (1.66 with clypeus), prosoma 0.57 long, 0.55 wide, opisthosoma 1.01 long, 0.53 wide. Legs I and II missing, leg III: 5.05 (1.37 + 0.19 + 1.23 + 1.74 + 0.52), leg IV: – (2.30 + 0.20 + 2.06 + – + –). Eye interdistances and diameters: PME-PME 0.14, PME 0.06, PME-ALE 0.03, AME absent. Sternum width/length: 0.45/0.45. Habitus as in Figs 5C–E . Dorsal shield of prosoma and sternum whitish, without marks. Legs whitish, without darker rings. Opisthosoma whitish, without spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae ( Fig. 5B ) with pair of proximo-lateral apophyses and pair of curved distal apophyses (distance between tips: 0.29). Pedipalp as in Figs 4A–B ; trochanter as long as wide; femur with retrolatero-proximal apophysis (arrow in Fig. 4B ); procursus ( Figs 4A–D , 9C–D ) simple proximally but complex distally, with prolatero-distal membranous lamella with weakly sclerotized lamella (arrow 1 in Figs 4C , 9C ), two spine-shaped dorso-distal apophyses (arrows 2, 3 in Figs 4C , 9C ), ventral membranous lamella (arrow 4 in Figs 4C , 9C ), retrolatero-distal membranous lamella (arrow in Figs 4D , 9D ), and retrolatero-subdistal membranous flap (f in Figs 4D , 9D ); bulb ( Fig. 5A ) with hooked apophysis and simple embolus. Legs with short vertical setae on metatarsi.

Female: Unknown.

Natural history. The species was collected in a seasonal rainforest by canopy fogging.

Distribution. China (Yunnan, type locality; Fig. 1 ).

Figures

FIGURE 1. Distribution records of the Belisana species discussed in this paper. 1, B. chuandiani sp. nov.; 2, B. daxiangi sp. nov.; 3, B. fengzheni sp. nov.; 4–5, B. zhengi.

FIGURE 4. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310). A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.02 (C–D).

FIGURE 5. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310).A. Bulb, prolateral view; B. Chelicerae, frontal view; C–E. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.03 (A–B), 0.20 (C–E).

FIGURE 6. Belisana fengzheni sp. nov., holotype male (SYNU-Ar00311). A–B. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); C–D. Distal part of procursus (C. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.02 (C–D).

FIGURE 7. Belisana fengzheni sp. nov., holotype male (C, E–F; SYNU-Ar00311), paratype male (D, SYNU-Ar00312) and paratype female (A–B, G–H; SYNU-Ar00315). A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.20 (E–H).

FIGURE 9. Procursus in prolateral and retrolateral views.A–B. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); C–D. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); E–F. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10.

FIGURE 10. Female genitalia in ventral and dorsal views. A–B. Belisana chuandiani sp. nov.; C–D. Belisana fengzheni sp. nov.; E–F. Belisana zhengi Yao, Pham & Li, 2015. ep = epigynal pocket, pp = pore plate. Scale bars: 0.10.