Meoneura simplex, Papp, 2013

Papp, L., 2013, New Species And Records Of Meoneura Rondani, 1856 From The Old World Tropics (Diptera, Carnidae), Acta Zoologica Academiae Scientiarum Hungaricae 59 (3), pp. 213-227 : 219-221

Meoneura simplex sp. n.

(Figs 16–20)

Holotype male ( HNHM): India, Uttar Pradesh, Nainital, Governor’s House Park , ca. 2050 m – swept and singled, 7. XII. 1989, leg. Papp L.

Paratypes ( HNHM): 2 males 1 female (1 m with genitalia preparation): same data ; 1 male: ibid., near Tiffin Top, 2100 m – swept and singled, 5. XII. 1989 .

Measurements in mm: body length 1.47 (holotype), 1.43–1.70 (paratypes), wing length 1.45 (holotype), 1.29–1.80 (paratypes), wing width 0.60, 0.55–0.69.

Body subshiny black, legs dull black.

Frons almost all black, only most anterior margin reddish on c. 0.05 mm length. Frontal triangle broad, shiny, reaching 3/7–4/9 distance of anterior ocellus to frontal margin. Face dark grey. Supralunular (interfrontal seta) 0.12–0.13 mm long. Outer and inner occipital setae short. Dorsal anterior (actually cheek) genal seta 0.13 mm, ventral anterior genal seta 0.13 mm, seta between them half as long; 2 similarly long posterior genal seta on middle of gena, one of them strongly upcurved. Ventral genal setae short. Gena anteriorly (at narrowest) 0.10 mm. Eye 0.275 × 0.25 mm. Arista 0.21 mm long.

Mesonotum black, thinly grey microtomentose. Anterior 2 dorsocentral pairs shorter, the posterior dc long, anterior dc just on suture. Longest anepisternal seta 0.14–0.15 mm, longest katepisternal seta 0.14 mm long.

Legs all microtomentose black. Fore femur with 3 long posteroventral setae.

Figs 16–25. Meoneura species , male genitalia: 16–20 = M. simplex sp. n., paratype male: 16 = contours of epandrium, cerci and subepandrial sclerite, caudal view (subepandrial sclerite dashed), 17 = surstylus in broadest extension, sublateral view, 18 = postgonite, lateral view, 19 = same, subanterior view, 20 = same, anterior view. 21–25 = M. subinversa sp. n., para- type male: 21 = left surstylus in broadest extension, sublateral view, 22 = right surstylus in broadest inner (medial) view, 23 = postgonite, lateral view, 24 = postgonite, caudal view,

25 = lamella, broadest inner (a sublateral-subcaudal) view. Scales: 0.1 mm for Fig. 21, and

16–20, 22–25, respectively.

Wing membrane pale grey, veins (incl. costal vein) yellow. Interfrontal section of M 1+2 0.08 mm, M-M crossvein 0.04 mm long. Alula narrowly triangular. Knob of haltere white, stalk light brownish.

Abdominal sternite 1 well-formed, transverse, sternite 2 longer than broad, sternite 3 and 4 more than twice longer than broad, male sternite 5 0.10 mm long and 0.10 mm broad, marginal setae short, only 0.03 mm. Pregenital sclerite 0.24 mm broad but only 0.09–0.10 mm long. Subepandrial sclerite not long, arched (Fig. 16), epandrium forms a pair of subtriangular processes below cerci in contrast to M. hungarica L. Papp ( PAPP 1977: fig 8, 1978: fig. 28A). Epandrium with several strong setae in distal (caudal) half, longest seta 0.09 mm. Surstylus (Fig. 17) broad based, strongly narrowing distally but less so than in M. hungarica , apex more broadly rounded (cf. PAPP 1977: fig. 8). No lamella developed at all. Postgonite (Figs 18–20) very characteristic: short and broad-based in lateral view (Fig. 18) with short proclinate apex; comparison of its lateral and anterior views though a subanterior view (Fig. 19) shows that apex is also outward directed (Fig. 20). Phallus similar to that of M. indica sp. n., apical 2/3 strongly swollen and turns to the right.

Female without peculiarities, so not safely identifiable in lack of synchronously cap- tured males.

Etymology. The specific epithet of the new species ‘simplex’ refers to its very simple genitalia.

I did not find any closer relative than M. triangularis Collin, 1930 (a Holarctic species). However, the surstylus of M. triangularis is really triangular with acute though not sharp apex. Setae on caudal sub-basal part of surstylus seem somewhat longer and thicker than in M. triangularis . Females cannot be safely separated.