Polynema (Polynema) grafinya Triapitsyn, 2021

Triapitsyn, Serguei V., 2021, Taxonomic review of Polynema (Hymenoptera: Mymaridae) in Australia and New Zealand, with descriptions of eight new species, Zootaxa 4915 (2), pp. 151-200 : 178-180

publication ID	https://doi.org/ 10.11646/zootaxa.4915.2.1
publication LSID	lsid:zoobank.org:pub:6E0C3BA7-7C8B-4D80-AFE4-07AEC6DEC71C
DOI	https://doi.org/10.5281/zenodo.4454230
persistent identifier	https://treatment.plazi.org/id/03C9DE3E-FFF9-FFF6-0DA6-FC3DFD64F891
treatment provided by	Plazi
scientific name	Polynema (Polynema) grafinya Triapitsyn
status	sp. nov.

Treatment

Polynema (Polynema) grafinya Triapitsyn , sp. n.

( Figs 73–82 View FIGURES 73–78 View FIGURE 79 View FIGURES 80–82 )

Type material. Holotype female [ NZAC] on slide ( Fig. 74 View FIGURES 73–78 ) labeled: 1. “ NEW ZEALAND: MB, Red Hills, Wairau, 22 Mar. 1972, E. W. Valentine, E1121/4 ENTOMOLOGY DIVISION D.Z.I. R. NZ”; 2. “ ♀ Polynema 28 NOV 1980 ”. The type locality is on South Island.

Paratypes: NEW ZEALAND. South Island : Coronet Peak (OL), 1640 m, i.1981, J.S. Noyes, E.W. Valentine (swept, Tussock/Alpine shrubs, Hebe , mat plants) [1♀ on slide, NZAC]. Wairau ( MB), Red Hills, 22.iii.1972, E.W. Valentine (E1121) [11♀ on 3 cards (6, 3, and 2 respectively), 1Ƌ on slide, NZAC] .

Description. FEMALE (holotype). Body and appendages dark brown except petiole and legs (the latter partially) brown (as in Fig. 79 View FIGURE 79 ). Head ( Fig. 76 View FIGURES 73–78 ) wider than high; face with fine, inconspicuous subtorular lines. Antenna ( Fig. 73 View FIGURES 73–78 ) with scape, including short radicle, 3.3× as long as wide, smooth; pedicel smooth, longer than F1 and 2.0× as long as wide; F1 the shortest and F2 the longest funiculars, F3 as long as F6 and longer than F4 or F5, length to width ratios of funiculars: F1—2.3, F2—6.1; F3—4.2; F4—2.4; F5—2.2; F6—2.6, F6 with 1 mps; clava 2.6× as long as wide, about as long as combined length of 3 preceding flagellomeres, with 7 mps. Mesosoma about 1.7× as long as wide, mostly smooth except mesoscutum with fine, inconspicuous mesh-like sculpture; pronotum entire; mesoscutum wider than long, with notaular grooves; scutellum shorter than mesoscutum, with campaniform sensilla closer to anterior than to posterior margin and widely separated from each other but at a distance from lateral margins (as in Fig. 81 View FIGURES 80–82 ), and with frenal row of small foveae; propodeum ( Fig. 75 View FIGURES 73–78 ) with a median carina extending from posterior margin for about half length. Fore wing ( Fig. 78 View FIGURES 73–78 ) 5.0× as long as wide; marginal vein with 1 dorsal macrochaeta; disc slightly infumate, densely setose beyond venation; the discal setae originating behind apex of submarginal vein; longest marginal seta 1.0× greatest width of wing. Hind wing about 33× as long as wide; disc slightly infumate, with an incomplete median row of setae; longest marginal seta 5.3× greatest width of wing. Metacoxa almost smooth. Petiole with anterolateral “wings”. Ovipositor occupying almost entire length of gaster ( Fig. 77 View FIGURES 73–78 ), exserted beyond gastral apex by 0.14× own total length, and 1.7× length of metatibia.

Measurements of the holotype (µm). Mesosoma: 320; gaster: 449; ovipositor: 486. Scape: 121; pedicel: 60; F1: 30; F2: 85; F3: 55; F4: 38; F5: 41; F6: 55; clava: 136. Fore wing: 898:179; longest marginal seta: 182. Hind wing: 763:23; longest marginal seta: 121.

Variation (paratypes). Body length (dry-mounted specimens, as in Fig. 79 View FIGURE 79 ) 825–960 µm. In the slide-mounted specimen, scape, including radicle, 3.5× as long as wide, clava 2.3× as long as wide, fore wing 5.5× as long as wide, longest marginal seta 1.3× greatest width of fore wing, and ovipositor exserted beyond gastral apex by about 0.1× own total length, and 1.6× length of metatibia.

MALE (paratype). Similar to female in color except petiole ( Fig. 81 View FIGURES 80–82 ) brown. Antenna ( Fig. 80 View FIGURES 80–82 ) with scape, excluding radicle, 2.2× long as wide and shorter than any funicular. Fore wing ( Fig. 82 View FIGURES 80–82 ) 5.4× as long as wide; longest marginal seta 1.1× greatest width of wing. Genitalia typical for species of P. ( Polynema ) ( Triapitsyn & Fidalgo 2006), 230 µm long; digitus with 2 small denticles apically .

Diagnosis. Polynema grafinya females can be separated from females of other fully winged species of the draperi group by its relatively long ovipositor, at least 1.6× the length of the metatibia, that occupies almost the entire length of the gaster and is exserted beyond the gastral apex by at least 0.1× its own total length ( Fig. 77 View FIGURES 73–78 ).

Etymology. The species name is a noun in apposition; it means countess in Russian.

Remarks. Three females [NZAC] from South Island, New Zealand, labeled: “ NEW ZEALAND, OL, Kinloch SF, Dart R, Jan. 1981, Nothofagus For Broadleaf , grass P. totara Swept, J. S. Noyes, E. W. Valentine ” may also belong to this species; however, they have a relatively narrower fore wing (5.7–6.0× as long as wide) and a shorter ovipositor (about 1.5× as long as metatibia).