PULMONATA

Dayrat, B. & Tillier, S., 2002, Evolutionary relationships of euthyneuran gastropods (Mollusca): a cladistic re-evaluation of morphological characters, Zoological Journal of the Linnean Society 135 (4), pp. 403-470 : 461

PULMONATA, STYLOMMATOPHORA

Succinea Draparnaud, 1801 (Baudon, 1877; Schnabel, 1903; Quick, 1933, 1939; Franzen, 1959; Duncan, 1961; Rigby, 1965; Cook, 1966; Delhaye & Bouillon, 1972c; Bayne, 1973; Solem, 1976; Visser, 1977, 1981). (43–51) We code the visceral loop as {(CPl)(SbAbSp)(PlC)} (Cook, 1966). (57) The albumen gland is ambiguous (Rigby, 1965).

Achatinella Swainson, 1828 (Pilsbry, 1900; Cooke & Kondo, 1960; Van Mol, 1967; Tillier, 1984, 1989). (32) The ureter of orthurethrans is interpreted either as secondary (Cooke & Kondo, 1960) or as primary (Delhaye & Bouillon, 1972c). (43–51) We code the visceral loop as {PlSb(AbSp)Pl} (Van Mol, 1967; Tillier, 1984, 1989). (57–60) The species of the genus Achatinella are ovoviviparous: eggs are stored in the oviduct and nidamental glands are consequently absent.

Achatina Lamarck, 1799 (Reynell, 1906; Mead, 1950; Ghose, 1962a, b, c, 1963a, b, c, d, e, 1964; Hyman, 1967; Van Mol, 1967; Brisson, 1968; Breckenbridge & Fallil, 1973; Tillier, 1989). (1) The embryological shell is dextral but seems to be senestral at the begining of development (Brisson, 1968). (25) A larval kidney (Brisson, 1968) is probably homologous with the PMO. (43–51) The topology of the visceral loop {PlPaAbPaPl} (Ghose, 1962; Tillier, 1989) is interpreted here as {PlSbAbSpPl}.

Arion Férussac, 1819 (Lamy, 1929; Van Mol, 1962; Smith, 1966; Van Mol et al., 1970; Parivar, 1978). (43–51) We code the visceral loop as {(PlSb)Ab(SpPl)}, {Pl(SbAbSp)Pl} or {(PlSb)(AbSp)Pl} (Van Mol, 1962).

Helix Linné, 1758 (Schnabel, 1903; Prenant, 1924, 1926; Lamy, 1929; Duncan, 1960a; Lind, 1973; Kerkut & Walker, 1975; Visser, 1977; Godan, 1979; Enée & Griffond, 1983; Tillier, 1989). (1) The embryological shell is dextral (Godan, 1979). (43–51) We code the visceral loop as {PlSbAbSpPl} (Kerkut & Walker, 1975; Tillier, 1989). No parietal ganglia appear during development (Fol, 1879).