Cydistomyia madangiensis, Goodwin, 2010

Goodwin, James T., 2010, New species of Cydistomyia Taylor with notes and collection records for other Tabanidae (Diptera) from New Guinea, Insecta Mundi 2010 (117), pp. 1-22 : 5-6

Cydistomyia madangiensis n. sp. ( Figure 2 A-2D)

Material examined. 3 females. Holotype female: on leaf {of} Theobroma cacao , near Sapi Village, Madang Province, PNG, 1-VI-1984, by H. Roberts ( FSCA; D1454); paratype female: Castanopsis Bamboo Forest / Manki L.A. rd., Bulolo, M. Prov., 14 XI-1983, J. Dobunaba ( FSCA; D1455); paratype female: NEW GUINEA: NE, Wau, 1100-1200m, VI (June) 1968, N. H. L. Krause ( BPMB).

Description. A medium-sized (body and wing lengths 12 mm) brown species lacking distinct pattern except for pale hind margins on abdominal segments 2-6.

Head. Frons ca. 0.6 mm wide basally and ca. 5 times as high as wide basally, about same width above and below but distinctly narrowed in middle, yellowish pollinose overall but slightly darker over middle of upper one-half; basal callus pale brown, large, narrowly separated from eyes, about 2.5 times as high as wide basally, narrowing into an elongate median callus that tapers to a pointed apex over upper half of its length. Subcallus, parafacials, and face yellowish-gray pollinose, subcallus a little darker over middle one-third. Antennae orange-brown with abundant black hairs on scape and pedicel and a few on the tooth of the basal plate; basal plate with a shallow dorsal excision, about 1.5 times as long as broad, slightly longer than combined length of annuli (style). Palpi rather stout and short, brown, faintly grayish dusted and covered by abundant black hairs.

Thorax. Mesonotum and scutellum grayish pollinose over brown base color, the scutellum a little lighter in color, lacking pattern. Pleural areas grayish pollinose with white hairs. Legs with coxae gray- ish pollinose with white hairs, remainder pale brown with mostly black hairs, the tarsi a little darker. Wings clear except for pale yellowish tint of costal cell.

Abdomen. Pale brown dorsally on tergites 1 and 2, darker brown on remainder; posterior margins of tergites 2-6 with narrow grayish pollinose margins. Venter blackish-brown with pale hind margins on sternites 2-6.

Etymology. The specific epithet is based on the name of the province in which the holotype was collected.

Discussion. This species would be part of the immigrans group ( Mackerras, 1964, 1971) recognized by Mackerras (1964, 1971). In Mackerras (1964) it would most likely key to C. torresi from which it differs as follows: in C. madangi the callus is wedge-shaped, tapering gradually to a pointed apex that ends just below middle of frons, the frons diverges slightly from near its middle to the union with the subcallus, and the subcallus is grayish pollinose as are the parafacials and face; in C. torresi , the frons is parallel sided and 5 or more times as high as wide, the callus has an expanded base that is taller than wide and has a slender median dorsal extension that ends distinctly above the middle of the frons, and the subcallus is brown, contrasting with the grayish white parafacials and face.

Figures

Figure 2. Adult habitus and structures of Cydistomyia madangi n. sp. A) Outline drawing of antenna. B) Outline drawing of palp. C) Anterior view of head. D) Dorsal habitus. Scale line 0.1mm.