Anopinella cartagoa Brown and Adamski

Brown, John W. & Adamski, David, 2003, Systematic revision of Anopinella Powell (Lepidoptera: Tortricidae: Euliini) and phylogenetic analysis of the Apolychrosis group of genera, Zootaxa 200, pp. 1-94 : 14-16

Anopinella cartagoa Brown and Adamski , new species

Figs. 4 , 37 , 59

Diagnosis. Anopinella cartagoa is a moderately large species, superficially most similar to A. triquetra and A. porrasa . The male genitalia are easily distinguished by the presence of a keel­like process from the venter of the subapical portion of each of the gnathos arms and a long, strong uncus, both of which appear to represent autapomorphies for the species. The female genitalia are distinguished by the anteriorly convergent, sclerotized ridges of the eighth tergum, with a short, mesal, digitate lobe at the anterior edge, and the shape of the ductus bursae: broad at the antrum and gradually tapered anterad.

Description. Head: Frontoclypeus and vertex pale brown intermixed with brown; labial palpus with outer surface of segments I­II brown intermixed with pale brown, segment III pale brown, inner surface pale brown. Antenna with scape pale brown intermixed with brown; basal 8­10 flagellomeres reddish brown; distal flagellomeres gray.

Thorax: Tegula and mesonotum pale brown intermixed with brown and dark brown. Forewing (Fig. 59) length 7.7­8.8 mm (n = 10); incomplete basal fascia and costal blotch brown intermixed with reddish brown and dark brown, separated by an oblique, pale band of white intermixed with pale brown and reddish brown demarcating anterior part of basal fascia and costa blotch to slightly beyond CuP, recurved from distal costa through subapical area to tornus, nearly entirely encircling an elongate ocellus, except for posterior part; ocellus rounded apically, wide, posteriorly narrowed or truncate, pale brown intermixed with pale reddish brown and white; costal blotch with a small subrectangular white spot near posterior end; area between CuP and posterior margin reddish brown intermixed with brown; apical and submarginal areas narrow, brown intermixed with reddish brown. Fringe brown intermixed with reddish brown. Hindwing pale grayish brown.

Abdomen: Male genitalia ( Fig. 4 ; drawn from DA slide 4746; n = 4) with uncus long, large, broadly curved throughout. Socius narrow, digitate, short, ca. 0.17 length of gnathos arms. Gnathos relatively wide, gradually narrowed distally; distal lobes slightly upturned, acuminate, with entire margins, dorsally connected by a wide hood­shaped lobe; ventral margin with a small, semicircular, keel­like, subapical process. Valva gradually widened from subbasal curve, densely setose from inner surface of cucullus to basal ridge; costa slightly arched from subbasal curve; ventral margin broadly rounded, slightly recurved at apicoventral angle; cucullus slightly acuminate, with outer margin nearly straight from ca. 0.33 length to apicoventral angle. Phallus simple; vesica sparsely microtrichiate. Female genitalia ( Fig. 37 ; drawn from DA slide 4560; n = 12) with weakly developed subquadrate sclerotized mesal patch at lamella postvaginalis; eighth tergum with a pair of oblique, sclerotized lines (representing flattened lateral lobes at the anterior end of segment) terminating in a short, mesal, digitate lobe anteriorly ( Fig. 37 insert); ostium wide, shallowly Ushaped, ca. 0.33 width of seventh sternum at widest point. Ductus bursae long, moderately broad at antrum, gradually tapered anterad; inception of accessory bursae ca. 0.8 distance from ostium to corpus bursae. Corpus bursae ovoid, spiculate on ca. 0.2 of side bearing ductus seminalis.

Holotype, ɗ, Costa Rica , Cartago Province, Paraíso, P.N. Tapantí­Macizo de la Muerte, Al Costado de Casa Adm., 1200 m, Jun 2000, L. Chavarria. Deposited in INBio.

Paratypes (3ɗ, 14Ψ). COSTA RICA : Cartago Province: Paraíso, P.N. Tapantí­Macizo de la Muerte, Al Costado de Casa Adm., 1300 m, Jun 2000 (1ɗ), R. Delgado (INBio). Paraíso, P.N. Tapantí­Macizo de la Muerte, 300 m SE del Puente del Río Porras, 1660 m, Jan 2000 (1Ψ), Nov 2001 (1ɗ, 4Ψ), Aug 2002 (1Ψ), R. Delgado (INBio, USNM); Paraíso, P.N. Tapantí­Macizo de la Muerte, Estación Quebrada Segundo, 1300 m, Mar 2001 (1Ψ), Aug 2000 (1Ψ), R. Delgado (INBio). Tapantí, 1700 m, 10 Jun 1998 (1Ψ), 1500 m, 30­31 Aug 2000 (1Ψ), V. O. Becker (VBC). Guanacaste Province: Z. P. Tenorio, Sector Alto Masís, 1100 m, 10­14 Jun 2002 (1Ψ), J. Jiménez (INBio), 10­14 Jun 2002 (1Ψ), L. Chavarria (INBio). Heredia Province: 16 km SSE La Virgen, 10 16'N, 84 05'N, 1050­1150 m, 19 Feb 2001 (1ɗ), D. Wagner & J. Rota (INBio), 19 Apr 2001 (1Ψ), J. Brown (USNM). Puntarenas Province: Buenos Aires, PILA, Sector Altamira, A.C Amistad, 1150­1400 m, May 1994 (1Ψ), R. Delgado (INBio).

Etymology. The specific epithet is derived from Cartago Province, Costa Rica .

Remarks. The association of the sexes is based on the series taken at Paraíso in November 2001, which includes both males and females. Although the holotype of A. porrasa was collected at the same locality, the latter has a conspicuously longer forewing length.

Figures

FIGURES 3 ­ 4. Male genitalia of Anopinella, valvae spread, aedeagus removed. 3, A. ophiodes; 4, A. cartagoa.

FIGURES 36 ­ 37. Female genitalia of Anopinella, posterior end up. 36, A. porrasa, with insert of tergum eigth; 37, A. cartagoa, with insert of tergum eight.