Diplommatina fistulata Budha & Naggs, 2017

Budha, Prem B., Naggs, Fred & Backeljau, Thierry, 2017, The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species, European Journal of Taxonomy 337, pp. 1-30 : 8-9

Diplommatina fistulata Budha & Naggs sp. nov.

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Figs 3C , 9C

Diagnosis

Measurements (n = 10): SH 3.0– 3.3 mm, SW 1.5–1.8 mm, Wh 6.5–7.5; holotype: SH 3.3 mm, SW 2.0 mm, Wh 7.5. Dextral, body whorl narrower than penultimate whorl. Constriction at the middle of the parietal lip ( Fig. 9C ). Parietalis long and very strong ( Fig. 3C3 ). Two vertical palatalis: a stronger Vpt1 along the constriction and a short Vpt2 running parallel to Vpt1 above the parietal wall ( Fig. 9C ). No horizontal palatalis. Columellaris prominent. Columellar tooth strong. Protoconch smooth, Wh 2.0, with minute pits scattered over the surface ( Fig. 3C 1 ). Teleoconch with dense radial ribs; 9–11 ribs/ 0.5 mm on the penultimate whorl; 15–20 ribs/ 0.5 mm on the body whorl towards the aperture ( Fig. 3C 2 , C 4 ). Distinct dorsal fold at base of body whorl, running towards the aperture ( Fig. 3C 4 ). Spiral striation present. Umbilical perforation visible, but umbilicus seems to be closed by the reflected edge of the columellar lip. Aperture ear-shaped, parietal wall oblique with regard to the coiling axis; peristome with two close lips; inner lip thickened and forming a thick callus over the parietal wall; under high magnification 12–15 highly dense ribs visible between the lips. Operculum paucispiral, corneous, thin, outer surface smooth, inner surface with thick peripheral circular edge.

Etymology

The species name refers to the tube-like structure of the dorsal fold on the body whorl just behind the aperture.

Type material

Holotype

NEPAL: Phulchowki Hill , Lalitpur District, 2308–2483 m, 27.574557 N, 85.400842 E, 4 May 2007, P.B. Budha leg. ( CDZMTU025 ).

Paratypes

NEPAL: 5 shells from the type locality, same data as holotype ( CDZMTU 026); 4 shells, Baghdwar, Shivapuri-Nagarjun National Park, 2326 m, 27.8131 N, 85.4217 E, 22 Jun. 2007, P.B. Budha leg. ( CDZMTU 027a); 8 shells, Baghdwar area, Shivapuri-Nagarjun National Park, 2412–2502 m, 27.8233 N, 85.3877 E, 24–25 Apr. 2008, R. Devkota and P.B. Budha leg. ( CDZMTU 027b); 2 shells, Dhunche, Langtang National Park, 2056 m, 28.1087 N, 85.2916 E, 3–4 Jun. 2007, P.B. Budha leg. ( CDZMTU 057).

Remarks

Diplommatina fistulata sp. nov. resembles the NE Indian D. pachycheilus , from which it differs by its distinct dorsal fold behind the aperture, smaller size and more slender shape.

Figures

Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm.

Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov.

Fig.1. Map of the localities and areas surveyed for Diplommatina spp. in Nepal.

Fig. 2. Shell terminology. A. D. regularis Fulton, 1901. B. D. folliculus (L. Pfeiffer, 1846). C. General outline of the lower part of a shell from Diplommatina species. Abbreviations: Prt = Parietalis (Prt1 = inner parietalis, Prt2 = outer parietalis); Pt = Palatalis; Hpt = Horizontal palatalis (Hpt1 = lower, Hpt2 = upper in case of two); Vpt = Vertical palatalis (Vpt1 = left side, Vpt2 = right side in case of two).

Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm.