Goja tamensis, Carrejo & Obando & Aldrete & Mendivil, 2021

Carrejo, Nancy, Obando, Ranulfo González, Aldrete, Alfonso Neri García & Mendivil, Julián, 2021, New species of Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) from Colombia Mexico and Peru, Zootaxa 4903 (4), pp. 501-541 : 532

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Goja tamensis

n. sp.

Goja tamensis n. sp. Male

( Figs 100–105 View FIGURES 100‒105 )

Diagnosis. Forewing R2+3 dichotomously branched, R4+5 two-branched, the second branch forked as illustrated, resulting in three veins at wing margin; a crossvein between R2+3 and R4+5. M six-branched, with M1 and M2 forked. Hindwing Rs and M three branched. Anterior endophallic sclerite a transverse, almost semicircular body bearing posteriorly two short, acuminate, curved apophyses on each side of the midline; posterior endophallic sclerites each constituted by a large, almost elliptical body, bearing posteriorly a slender, curved outward process, acuminate distally ( Fig. 105 View FIGURES 100‒105 ). It is similar to G. circumvittata Casasola González & García Aldrete , from which it differs in forewing venation and endophallic sclerites.

Color (in 80% ethanol). Body brown. Head dark brown ( Fig. 102 View FIGURES 100‒105 ), color pattern as illustrated; striae of postclypeus tenuous. Compound eyes dark brown, ocelli hyaline, with ochre centripetal crescents. Maxillary palps pale brown, Mx4 dark brown distally. Antennae pale brown. Thorax brown, tergal lobes of meso- and metathorax dark brown, mesopleura brown. Legs pale brown, apices of femora and tarsi dark brown. Forewings ( Fig. 100 View FIGURES 100‒105 ), pale brown, pterostigma dark brown, a marginal brown band from R4+5 to wing base; veins brown. Hindwings ( Fig. 101 View FIGURES 100‒105 ), almost hyaline, veins brown. Abdomen brown. Clunium, hypandrium, epiproct and paraprocts brown, hypandrium dark brown on sides. Phallosome brown, with endophallic sclerites and aedeagal arch dark brown.

Morphology. As in diagnosis, plus the following: Head: H/MxW: 1.40; compound eyes large, H/d: 4.81; IO/ MxW: 0.80. Outer cusp of lacinial tips broad, with six denticles. Mx4/Mx2: 1.11. Forewings ( Fig. 100 View FIGURES 100‒105 ): FW/W: 2.66. Pterostigma: lp/wp: 3.92. Areola postica wide, rounded: al/ah: 2.67. Hindwings ( Fig. 101 View FIGURES 100‒105 ): HW/w: 3.14. Hypandrium ( Fig. 104 View FIGURES 100‒105 ) wide, posteriorly rounded, setose. Phallosome ( Fig. 105 View FIGURES 100‒105 ) with side struts straight, slender; aedeagal arch V-shaped, with rounded apex; external parameres dilated, apically truncate and with short internal process, bearing pores posteriorly. Paraprocts ( Fig. 103 View FIGURES 100‒105 ), oval, with a macrosetae near sensory field, and a distal setal field; sensory fields with 23 trichobothria on basal rosettes. Epiproct ( Fig. 103 View FIGURES 100‒105 ) wide, convex anteriorly, rounded posteriorly; a setal field posteriorly, one macroseta mesally, other setae as illustrated.

Measurements. FW: 3725, HW: 2825, F: 950, T: 1560, t1: 635, t2: 230, ctt1: 23, f1: 830, f2: 730, f3: 510, f4: 340, f5: 180, f6: 180, f7: 150, f8: 145, f9: 140, f10: 120, f11: 160, Mx4: 267, IO: 440, D: 225, d: 160, IO/d: 2.75, PO: 0.71.

Specimens studied. Holotype male. COLOMBIA. Norte de Santander, National Natural Park Tamá, La Camacha, Pozo Negro, 7°21’N: 72°28’W, 2203 m., 21–29.XI.2003, Malaise trap GoogleMaps . Paratypes: 3 males, NNP Tamá, Toledito, 7°20’N: 72°29’W, 1971 m. 1 male, NNP Tamá, 7°25’N: 2262 m., 3.XI.2003; 1 male, 6.VIII.2005. 1 male NNP Tamá, Arenal Trail, 7°25’26.9”N: 72°26’31.8”W, 2541 m., 2.VII.2016, N. Carrejo. 1 male, NNP Tamá, La Colorada Trail, 7°27’5.1”N: 72°26’40.4”W, 2245 m., 3.VII.2016. R GoogleMaps . González. Led light trap.

Etymology. The specific epithet is a name in apposition that refers to the National Natural Park Tamá, Norte de Santander department, where the specimens were found.


Departamento de Geologia, Universidad de Chile