Goja camachensis, Carrejo & Obando & Aldrete & Mendivil, 2021

Carrejo, Nancy, Obando, Ranulfo González, Aldrete, Alfonso Neri García & Mendivil, Julián, 2021, New species of Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) from Colombia Mexico and Peru, Zootaxa 4903 (4), pp. 501-541 : 505-508

Goja camachensis n. sp. Male.

( Figs 7–11 )

Diagnosis. Forewing R2+3 and R4+5 forked. M five-branched. Hindwing apparently with M three branched (not illustrated). Phallosome unique, anterior pair of endophallic sclerites transverse, both ends acuminate, with denticles on anterior border; posterior pair slender, curved, bulged anteriorly, distally acuminate ( Fig. 11 ). Similar to G. toleditensis n. sp., from which it differs in the forewing branches of Rs and M (M five branched in G. camachensis n. sp. vs. four branched in G. toleditensis n. sp.), the arms of the aedeagal arch are straight in G. camachensis n. sp. and curved in G. toleditensis n. sp. in addition, the shapes of the endophallic sclerites are quite different in the two species.

Color (in 80% ethanol). Body creamy, with brown spots. Head pale brown ( Fig. 8 ), with pigmented pattern as illustrated, striae of postclypeus tenuous. Compound eyes dark brown, ocelli hyaline, with ochre centripetal crescents. Maxillary palps pale brown. Antennae pale brown. Thorax pale brown, pronotum cream, with brown spots; tergal lobes of meso- and metathorax with brown spots, mesopleura cream, with ochre brown spot. Legs: coxae, trochanters and femora cream, coxae with brown spot basally, tibiae and tarsi pale brown, tibiae with cream areas, tarsi 1 cream basally. Forewings ( Fig. 7 ), hyaline, pterostigma with anterior and posterior pale brown bands, a submarginal brown spot from cell r5 cell to areola postica, vein ends with brown spot. Hindwings hyaline, veins pale brown. Abdomen cream, with ochre brown subcuticular spots. Clunium, epiproct, paraprocts and hypandrium brown, darker on sides; epiproct and paraprocts pale brown. Phallosome pale brown, endophallic sclerites, side struts and aedeagal arch dark brown.

Morphology. As in diagnosis, plus the following: Head: H/MxW: 1.70; compound eyes large, H/d: 4.22; IO/ MxW: 0.67. Outer cusp of lacinial tips broad, with two broad denticles, the most external being the widest. Mx4/ Mx2: 1.05. Forewings ( Fig. 7 ): FW/W: 2.74. Pterostigma long, widest in the middle: lp/wp: 6.42; areola postica low, elongate: al/ah: 4.26. Hindwings incomplete. Hypandrium ( Fig. 10 ), setose as illustrated, posterior border straight. Phallosome ( Fig. 11 ) with side struts slender, wide anteriorly, curved inward; aedeagal arch V-shaped, apex with short projection, rounded apically; external parameres rounded apically, bearing few pores on membranous area. Paraprocts ( Fig. 9 ), oval, with setae and macrosetae on distal setal field; sensory fields with 32 trichobothria on basal rosettes. Epiproct ( Fig. 9 ) wide, convex anteriorly, rounded posteriorly, with two lateral macrosetae, one macroseta mesally, posterior margin with setae as illustrated.

Measurements. FW: 5350, F: 1310, T: 2340, t1: 1020, t2: 270, ctt1: 38, Mx4: 305, IO: 380, D: 280, d: 230, IO/d: 1.65, PO: 0.82.

Specimens studied. Holotype male. COLOMBIA. Norte de Santander, Tamá National Natural Park , La Camacha , Pozo Negro , 7°21’N: 72°28’W, 2203 m., 21–29.XI.2003. 1 male, PBN Tamá , Toledito , 7°20’N: 72°29’W, 1970 m. IAvH-E 145913, code 3903. Malaise trap.

Etymology. The specific epithet refers to the type locality, village La Camacha.

Figures

FIGURES 7‒11. G. camachensis n. sp. Male. 7. Forewing. 8. Front view of head. 9. Left paraproct and epiproct. 10. Hypandrium. 11. Phallosome. Scales in mm.