Goja, Navás
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publication ID |
https://doi.org/ 10.11646/zootaxa.4903.4.3 |
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publication LSID |
lsid:zoobank.org:pub:CF96683E-75B8-4AC4-90A2-20AD8CC2F14B |
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DOI |
https://doi.org/10.5281/zenodo.4434603 |
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persistent identifier |
https://treatment.plazi.org/id/03C49710-FFEA-FFB1-598D-FBF7B78BFF0A |
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treatment provided by |
Plazi |
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scientific name |
Goja |
| status |
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Key to males of Goja View in CoL
(Modified from Casasola González & García Aldrete, 2002)
G. bimaculata , G. pallida and G. trinidadensis , all by Casasola González & García Aldrete, 2002, are not included because only females are known. G. camachensis n. sp., is not included because it lacks the distal part of the hindwing. G. poblana Casasola González & García Aldrete is not included because the male genitalia is missing.
1. Hindwing with caeciliusid (forewing Rs forked, M three branched; hindwing M simple) venation ( Figs 35 View FIGURES 34‒38 , 74 View FIGURES 73‒78 , 80 View FIGURES 79‒84 , 113 View FIGURES 112‒117 )... 2
- Hindwing Rs with two or more branches; M branched ( Figs 2 View FIGURES 1‒6 , 13 View FIGURES 12‒17 )............................................. 11
2. Forewing R4+5 two-three branched ( Fig. 73 View FIGURES 73‒78 )............................................................... 3
- Forewing R4+5 unbranched ( Figs 34 View FIGURES 34‒38 , 79 View FIGURES 79‒84 ).................................................................. 5
3. Forewing R4+5 distally forked; phallosome concave anteriorly, anterior endophallic sclerites U shaped, with inner arms straight, slender and acuminate; outer arms broad, ribbon-like, distally acuminate ( Fig. 131 View FIGURES 130‒141 )....................................................................................................... Goja moyai Casasola G. & García A. View in CoL
- Forewing R4+5 with three-four veins ( Figs 73 View FIGURES 73‒78 , 112 View FIGURES 112‒117 ); phallosome convex anteriorly, anterior endophallic sclerites not U shaped ( Figs 78 View FIGURES 73‒78 , 117 View FIGURES 112‒117 )........................................................................................ 4
4. Lateral endophallic sclerites with two posterior processes; outer processes acuminate, curved inward; inner processes distally rounded; aedeagal arch with apical process acuminate ( Fig. 78 View FIGURES 73‒78 )............................ Goja pillcopatensis View in CoL n. sp.
- Antero-lateral endophallic sclerites with two acuminate processes directed outward, mesal pair elongate and bent directed anteriorly, apices acuminate ( Fig. 117 View FIGURES 112‒117 ).................................................. Goja yarumosensis View in CoL n. sp.
5. Endophallic sclerites spiniform ( Fig. 38 View FIGURES 34‒38 ).................................................................. 6
- Endophallic sclerites U-shaped ( Fig. 120 View FIGURES 118‒129 )................................................................. 9
6. Head vertex well below the upper border of the compound eyes, these projected anteriorly ( Fig. 81 View FIGURES 79‒84 ); endophallic sclerites and aedeagal arch as in Fig. 84 View FIGURES 79‒84 ........................................................... Goja svanhildae View in CoL n. sp.
- Head vertex straight or slightly convex; compound eyes not projected as above ( Fig. 36 View FIGURES 34‒38 ); endophallic sclerites and aedeagal arch as in Fig. 38..................................................................................... 7 View FIGURES 34‒38 View FIGURES 7‒11
7. Apex of aedeagal arch pointed, with a distal process ( Fig. 128 View FIGURES 118‒129 )................. Goja mockfordi Casasola G. & García A. View in CoL
- Apex of aedeagal arch concave, with two distal process ( Fig. 38 View FIGURES 34‒38 )............................................... 8
8. Posterior endophallic sclerites stout, wide based, distally with three small lobes ( Fig. 138 View FIGURES 130‒141 )............................................................................................. Goja spinosissima Casasola G. & García A. View in CoL
- Posterior endophallic sclerites stout throughout, distally acuminate ( Fig. 38 View FIGURES 34‒38 )............... Goja cuasispinosissima View in CoL n. sp.
9. Genae unpigmented; apex of aedeagal arch projected ( Fig. 132 View FIGURES 130‒141 )............... Goja obrienorum Casasola G. & García A. View in CoL
- Genae with a pigmented area that reaches the antennal fossae; apex of aedeagal arch rounded ( Fig. 120 View FIGURES 118‒129 )............... 10
10. Anterior endophallic sclerites U-shaped, inner arms close together, about as long as the outer arms; epiproct with three mesal macrosetae....................................................... Goja brailovskyana Casasola G. & García A. View in CoL
- Anterior endophallic sclerites U-shaped, inner arms separated, shorter than the outer arms, these distally forked ( Fig. 120 View FIGURES 118‒129 ); epiproct with two mesal macrosetae.................................... Goja curvata Casasola G. & García Aldrete View in CoL
11. Hindwing R4+5 unbranched, M two branched ( Figs 51 View FIGURES 50‒55 , 57 View FIGURES 56‒61 , 68 View FIGURES 67‒72 )............................................... 12
- Hindwing R4+5 branched or not, M three-six branched ( Figs 13 View FIGURES 12‒17 , 45 View FIGURES 44‒49 )........................................... 24
12. Phallosome with side struts strongly curved; aedeagal arch extended posteriorly a distance, aedeagal apex broad and rounded; mesal sclerites elongate and curved, each one ended in an acute apex, joined to endophallus by long, twisted sclerotic bands......................................................................... Goja cuspidata Casasola González View in CoL
- Phallosome with side struts straight or not strongly curved as above; aedeagal arch not extended a distance posteriorly or apex not widened as above, mesal sclerites not as above ( Figs 105 View FIGURES 100‒105 , 111 View FIGURES 106‒111 )............................................. 13
13. Head without a pigmented spot posterior to antennal fossae.................................................. 14
- Head with a pigmented spot posterior or above the antennal fossae ( Fig. 108 View FIGURES 106‒111 ).................................... 19
14. Head with two ochre transverse bands, one between the compound eyes, through the ocellar triangle, and one between the genae, through the postclypeus ( Fig. 102 View FIGURES 100‒105 ); forewings R2+3 and R4+5 branched as illustrated ( Fig. 100 View FIGURES 100‒105 )............... 15
- Head without transverse bands as above ( Figs 52 View FIGURES 50‒55 , 69 View FIGURES 67‒72 ); forewing R2+3 forked or not, R4+5 forked or dichotomously branched ( Figs 50 View FIGURES 50‒55 , 67 View FIGURES 67‒72 )........................................................................................ 16
15. Forewings with R2+3 and Rs, after the R-M crossvein subequal; posterior endophallic sclerites robust, each with two small apical hooks; mesal endophallic sclerites each with a broad, longitudinal area, with a row of spines of unequal length on outer edge, directed outward, and a mesal spine on inner edge, directed anteriorly ( Fig. 141 View FIGURES 130‒141 )..................................................................................................... Goja vittata Casasola G. & García A. View in CoL
- Forewings with R2+3 almost twice as long as Rs after the R-M crossvein ( Fig. 94 View FIGURES 94‒99 ); posterior endophallic sclerites not as robust as above, apices rounded, without hooks; mesal endophallic sclerites each a broad, semioval area, without a row of spines as above ( Fig. 99 View FIGURES 94‒99 )..................................................................... Goja tacanaensis View in CoL n. sp.
16. Forewings with R2+3 as in Figs 50 View FIGURES 50‒55 and 67 View FIGURES 67‒72 ; phallosome as in Figs 55 View FIGURES 50‒55 and 72..................................... 17 View FIGURES 67‒72
- Forewings with R2+3 as in Fig. 56 View FIGURES 56‒61 ; phallosome variable ( Fig. 61 View FIGURES 56‒61 )............................................. 18
17. Mesal endophallic sclerites of three to four short acuminate teeth directed outward; lateral endophallic sclerites asymmetric, wide proximally, with distally acuminate elongate process curved outward ( Fig. 55 View FIGURES 50‒55 ); aedeagal arch V-shaped, apically acuminate............................................................................ Goja horquetensis View in CoL n. sp.
- Anterior pair of endophallic sclerites basally broad, irregular, each with a slender, long process on the inner side, directed posteriorly; posterior pair of endophallic sclerites well sclerotized, proximally wide, anterior border concave, with truncate process directed outward, distally with acuminate process curved outward ( Fig. 72 View FIGURES 67‒72 ); aedeagal arch rounded laterally, median process truncate........................................................................ Goja munchiquensis View in CoL n. sp.
18. Aedeagal arch robust, distally with a large, acuminate process; lateral endophallic sclerites broad, bent inward ( Fig. 124 View FIGURES 118‒129 )....................................................................... Goja chilensis Casasola G. & García A. View in CoL
- Aedeagal arch with posterior triangular process; lateral pair of endophallic sclerites kidney-shaped, with truncate posterior border; mesal pair with two elongate, acuminate processes directed outward ( Fig. 61 View FIGURES 56‒61 )............ Goja korytkowskii View in CoL n. sp.
19. Forewing R2+3 forked; R4+5 simple, a proximal crossvein between them ( Fig. 106 View FIGURES 106‒111 ); anterior pair of endophallic sclerites wide basally, semicircular, with posterior acuminate processes long and widely separated; posterior pair of endophallic sclerites oval ( Fig. 111 View FIGURES 106‒111 )......................................................................... Goja toleditensis View in CoL n. sp.
- Forewing R2+3 simple; phallosome variable.............................................................. 20
20. Forewing Rs about half as long as R2+3; lateral endophallic sclerites long, distally acuminate, each with two mesal spines directed outward, associated anteriorly to a transverse sclerite, each with an acuminate spine directed anteriorly ( Fig. 125 View FIGURES 118‒129 )................................................................... Goja longispinosa Casasola G. & García A. View in CoL
- Forewing Rs as long or longer than R2+3; phallosome with endophallic sclerites variable.......................... 21
21. Pigmented genal spots joined to the spots posterior to the antennal fossae; areola postica and M3 joined by a crossvein... 22
- Pigmented genal spots separated from the spots posterior to the antennal fossae; areola postica and M3 not joined by a crossvein.............................................................................................. 23
22. Apex of aedeagal arch with two acuminate distal processes; anterior pair of endophallic sclerites as vertical, membranous bands ( Fig. 118 View FIGURES 118‒129 )......................................................... Goja acuphallica Casasola G. & García A. View in CoL
- Apex of aedeagal arch with one rectangular distal process ( Fig. 133 View FIGURES 130‒141 ), anterior pair of endophallic sclerites with anterior processes stout, distally acuminate, posterior processes forked, robust, acuminate distally................................................................................................. Goja ochoterenae Casasola G. & García A. View in CoL
23. Apex of aedeagal arch wide, truncate, projected posteriorly ( Fig. 121 View FIGURES 118‒129 ), posterior endophallic sclerites wide at base, acuminate distally, each apex with two small hooks.................................. Goja eertmoedi Casasola G. & García A. View in CoL
- Apex of aedeagal arch rounded, not projected posteriorly ( Fig. 136 View FIGURES 130‒141 ), posterior endophallic sclerites slightly dilated anteriorly, slender, apices without small hooks.................................... Goja omiltemensis Casasola G. & García A. View in CoL
24. Forewings strongly pigmented, with large transverse hyaline areas; hindwing M three-five branched ( Figs 12 View FIGURES 12‒17 , 44 View FIGURES 44‒49 ); two pairs of endophallic sclerites, mesal pair broadly inverted U-shaped ( Figs 17 View FIGURES 12‒17 , 49 View FIGURES 44‒49 )........................................ 25
- Forewings unpigmented ( Fig. 23 View FIGURES 23‒27 ), if pigmented, then without hyaline areas as large as above; hindwing M three-four branched ( Fig. 24 View FIGURES 23‒27 ); phallosome variable.......................................................................... 26
25. Hindwing M five branched ( Fig. 13 View FIGURES 12‒17 ); anterior endophallic sclerites small, slender, distally acuminate, curved outward; mesal endophallic sclerites inverted U-shaped, with outer arms acuminate ( Fig. 17 View FIGURES 12‒17 )..................... Goja caucana View in CoL n. sp.
- Hindwing M three-four branched ( Fig. 45 View FIGURES 44‒49 ); mesal endophallic sclerites broadly inverted U-shaped, outer arms short, with a row of transverse lines along outer border; inner arms long, acuminate, slightly curved outward ( Fig. 49 View FIGURES 44‒49 )........................................................................................................ Goja galarzai View in CoL n. sp.
26. Forewing areola postica and M3 joined by one or several crossveins ( Fig. 23 View FIGURES 23‒27 ); hindwing R2+3 or R4+5 branched ( Fig. 24 View FIGURES 23‒27 ).. .................................................................................................. 27
- Forewing areola postica and M3 not joined by crossveins; hindwing R2+3 unbranched ( Figs 2 View FIGURES 1‒6 , 40 View FIGURES 39‒43 )................... 30
27. Forewing R2+3 without vein stubs; anterior endophallic sclerites broad, long; inner edge of each with a mesal, robust projection acuminate distally ( Fig. 139 View FIGURES 130‒141 )............................................... Goja ticana Casasola G. & García A. View in CoL
- Forewing R2+3 with vein stubs ( Fig. 23 View FIGURES 23‒27 ); anterior endophallic sclerites not as above ( Fig. 27 View FIGURES 23‒27 )....................... 28
28. Forewing R2 with one vein stub ( Fig. 23 View FIGURES 23‒27 ); aedeagal arch with short, narrow apical process ( Fig. 27 View FIGURES 23‒27 ); mesal endophallic sclerites with four pairs of spines, slender and distally acuminate, two anterior straight and directed outward, two short posterior pairs curved, acuminate............................................................... Goja chiquihuitensis View in CoL n. sp.
- Forewing Rs and R2+3 with several vein stubs; aedeagal arch apex wide, truncate, or blunt, spines of endophallic sclerites not as above........................................................................................... 29
29. Aedeagal arch apex wide, truncate, with two small distal processes; mesal endophallic sclerites robust, distally with one denticle ( Fig. 137 View FIGURES 130‒141 ), anterior pair slender, sinuous, wide basally, associated to a broad radula.................................................................................................. Goja reticulata Casasola G. & García A. View in CoL
- Aedeagal arch apex blunt, without terminal processes; mesal endophallic sclerites distally with two denticles, anterior pair broad, irregular, with acicular thickenings anteriorly...................... Goja reticulatoides Casasola G. & García A. View in CoL
30. Hindwing R4+5 unbranched ( Figs 2 View FIGURES 1‒6 , 40 View FIGURES 39‒43 )................................................................. 31
- Hindwing R4+5 branched ( Fig. 29 View FIGURES 28‒33 )...................................................................... 35
31. Hindwing R4+5 almost twice as long as Rs after the R-M crossvein................ Goja garcialdretei Casasola González View in CoL
- Hindwing R4+5 subequal in length to Rs after the R-M crossvein ( Figs 2 View FIGURES 1‒6 , 40 View FIGURES 39‒43 ).................................... 32
32. Aedeagal arch apex rounded, without processes ( Fig. 41 View FIGURES 39‒43 ), one pair of endophallic sclerites, anteriorly wide, elongate, acuminate apically and slightly curved inward...................................................... Goja farallones View in CoL n. sp.
- Aedeagal arch with apical processes pointed ( Fig. 6 View FIGURES 1‒6 ); endophallic sclerites variable................................ 33
33. Phallosome longer than wide, with four pairs of acuminate endophallic sclerites, anterior pair directed caudally and curved outward, the other three pairs directed outward; side struts straight ( Fig. 6 View FIGURES 1‒6 )..................... Goja caldasensis View in CoL n. sp.
- Phallosome almost as long as wide, with posterior endophallic sclerites caudally projected, side struts curved ( Figs 123, 126 View FIGURES 118‒129 ) .................................................................................................. 34
34. Apex of posterior endophallic sclerites acuminate; anterior endophallic sclerites horizontal, anteriorly wide, acuminate, directed outward ( Fig. 123 View FIGURES 118‒129 )............................................... Goja herrerae Casasola G. & García A. View in CoL
- Apex of posterior endophallic sclerites with two denticles ( Fig. 126 View FIGURES 118‒129 ), anterior endophallic sclerites broad, transverse, pointed at both ends, associated to two mesal ones, inverted Y-shaped.................. Goja martinezi Casasola G. & García A. View in CoL
35. Pterostigma without pigmented bands; anterior endophallic sclerites directed posteriorly, acuminate, each arm with six pointed processes directed outward ( Fig. 134 View FIGURES 130‒141 )..................................... Goja ocotensis Casasola G. & García A. View in CoL
- Pterostigma pigmented or with one-two pigmented bands ( Fig. 94 View FIGURES 94‒99 ); endophallic sclerites variable.................... 36
36. Pterostigma fully pigmented ( Fig. 100 View FIGURES 100‒105 ); anterior endophallic sclerites small, wide at base, acuminate distally, slender and curved outward ( Fig. 105 View FIGURES 100‒105 ).............................................................. Goja tamensis View in CoL n. sp.
- Pterostigma with pigmented bands, endophallic sclerites variable ( Fig. 28 View FIGURES 28‒33 )...................................... 37
37. Pterostigma with one pigmented band................................................................... 38
- Pterostigma with two pigmented bands ( Fig. 28 View FIGURES 28‒33 )........................................................... 39
38. Aedeagal arch apex blunt, without distal processes, posterior endophallic sclerites absent, anterior pair membranous ( Fig. 119 View FIGURES 118‒129 ) ….............................................................. Goja circumvittata Casasola G. & García A. View in CoL
- Aedeagal arch apex with an acuminate posterior process, and a blunt anterior process; anterior endophallic sclerites membranous, with two posterior, slender projections, on each side of mesal line ( Fig. 140 View FIGURES 130‒141 ).................................................................................................. Goja venezuelensis Casasola G. & García A. View in CoL
39. Forewing Rs, after R-M crossvein, one fourth or less the length of R2+3 ( Fig. 28 View FIGURES 28‒33 )................................. 40
- Forewing Rs, after R-M crossvein, about a third the length of R2+3............................................ 42
40. Aedeagal arch without apical process..................................... Goja medellini Casasola G. & García A. View in CoL
- Aedeagal arch with short apical process ( Fig. 33 View FIGURES 28‒33 ).......................................................... 41
41. Phallosome with mesal endophallic sclerites acuminate distally, curved back over slightly sclerotic area ( Fig. 122 View FIGURES 118‒129 )........................................................................ Goja guatemalensis Casasola G. & García A. View in CoL
- Phallosome with mesal endophallic sclerites directed forward, distally acuminate, not curved and directed backward as above ( Fig. 33 View FIGURES 28‒33 )................................................................... Goja cuasiguatemalensis View in CoL n. sp.
42. Forewing R4+5 two branched............................................................. Goja ditata Navás View in CoL
- Forewing R4+5 more than two branched................................................................. 43
43. Phallosome with posterior pair of endophallic sclerites more or less parallel, each with three digitiform small processes distally ( Fig. 129 View FIGURES 118‒129 ); hindwing M eight branched..................................... Goja montieli Casasola G. & García A. View in CoL
- Phallosome with posterior pair of endophallic sclerites diverging and acuminate distally ( Fig. 130 View FIGURES 130‒141 ); hindwing M four- five branched.......................................................................................... 44
44. Phallosome with posterior endophallic sclerites elongate and curved, each one ended in two conical acute apices, without spine basally ( Fig. 130 View FIGURES 130‒141 )......................................................... Goja bicuspidata Casasola González View in CoL
- Phallosome with posterior endophallic sclerites not as above, each with a stout anterior spine, postero-laterally directed, curved inward ( Fig. 135 View FIGURES 130‒141 )................................................... Goja oculatissima Casasola G. & García A. View in CoL
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.