Goja toleditensis, Carrejo & Obando & Aldrete & Mendivil, 2021

Carrejo, Nancy, Obando, Ranulfo González, Aldrete, Alfonso Neri García & Mendivil, Julián, 2021, New species of Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) from Colombia Mexico and Peru, Zootaxa 4903 (4), pp. 501-541 : 534

Goja toleditensis n. sp. Male

( Figs 106–111 )

Diagnosis. Forewing R2+3 forked, R4+5 unbranched, a crossvein between the two. M four branched, M3 and M4 distally forked. Hindwing venation caeciliusid. Anterior endophallic sclerites club-shaped, long, narrowing distally, acuminate, almost circular anteriorly: posterior endophallic sclerites large, almost circular. Similar to G. camachensis n. sp., for differences with it see the diagnosis of the latter above.

Color (in 80% ethanol). Body pale brown. Head cream ( Fig. 108 ), with ochre brown spots; genae cream, bottom with ochre spot. Compound eyes dark brown, ocelli hyaline, with ochre centripetal crescents; labrum brown, postclypeus with ochre striae. Thorax pale brown, tergal lobes of meso- and metathorax pale brown, pleura cream with ochre spots. Legs: mid- and hind- pale brown, mid- and hind- coxae with brown spots basally; femora with median and distal areas pale brown. Forewings ( Fig. 106 ), almost hyaline, veins brown, pterostigma with proximal and distal pale brown bands. Hindwings ( Fig. 107 ), hyaline, veins brown. Abdomen pale brown. Clunium, epiproct and paraprocts pale brown, hypandrium cream, posterior border and sides pale brown. Phallosome with aedeagal arch and endophallic sclerites brown, external parameres pale brown.

Morphology. As in diagnosis, plus the following: Head: H/MxW: 1.69; compound eyes large, H/d: 4.60; IO/ MxW: 0.67. Outer cusp of lacinial tips broad, without differentiated denticles. Forewings ( Fig. 106 ): FW/W: 2.61; pterostigma long: lp/wp: 5.77, narrow proximally; areola postica low, elongate: al/ah: 4.0. Hindwings ( Fig. 107 ): HW/w: 2.96. Hypandrium ( Fig. 110 ) concave posteriorly, a field of macrosetae on each side of the longitudinal midline, other setae as illustrated. Phallosome ( Fig. 111 ) bilobed anteriorly; side struts curved, slender, wide basally; aedeagal arch with short posterior process; external parameres well developed, joined to the side struts, wide basally, rounded posteriorly, with pores not clearly defined. Paraprocts ( Fig. 109 ), oval, with setae and macrosetae on distal field; sensory fields with 31 trichobothria on basal rosettes. Epiproct ( Fig. 109 ) wide, rounded posteriorly; setal field distally, three macrosetae mesally.

Measurements. FW: 4825, HW: 3625, F: 1205, T: 2080, t1: 730, t2: 250, ctt1: 29, IO: 366, D: 260, d: 200, IO/d: 1.83, PO: 0.77.

Specimens studied. Holotype male. COLOMBIA. Norte de Santander, National Natural Park Tamá, Toledito , 7°20’N: 72°29’W, 1970 m. Malaise trap. 1 male paratype, same data as the holotype.

Etymology. The specific epithet is a name in apposition that refers to the type locality, Toledito, were the specimens were found.

Figures

FIGURES 106‒111. G. toleditensis n. sp. Male. 106. Forewing. 107. Hindwing. 108. Front view of head. 109. Left paraproct and epiproct. 110. Hypandrium. 111. Phallosome. Scales in mm.