Gumilla Navás, 1912

Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian & Wang, Yongjie, 2019, Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera, Zootaxa 4581 (1), pp. 1-99 : 21

Gumilla Navás, 1912

( Figs 2C, J , 8–9 )

Type species. Gumilla adspersus Navás, 1912: 189 (by original designation).

Diagnosis. Antenna longer than FW with elongated flagellomeres and elongated scape; ocelli absent; wings ovoid; subcostal veinlets not forked; subcostal area with single sc-r crossvein; RP with crossveins irregularly arranged, without distinct gradate series; FW RP1 originating at approximately 1/3 of wing length, M fork from opposite to slightly beyond origin of RP1; HW medial fork at 2/3 of wing length from base to apex; male genitalia with gonarcus narrow with sparse pile; mediuncus (complex of gonocoxites 10 sensu Martins et al., 2016) narrow, arched, dorsomedially with a short subquadrate process; parameres (gonocoxites 11 sensu Martins et al., 2016) narrow, slightly arched; female genitalia unknown.

Comments. Navás (1912) correctly placed G. adspersus in Osmylidae but a reference by Hagen (1861) to Osmylus longicornis as a chrysopid based on the elongate antennae proliferated in the literature (e.g., Banks, 1907); the issue was settled by Adams (1977) who affirmed the conclusions of Navás (1912) and Krüger (1913b) that the latter species belonged in Gumilla . Two extant species in this genus are known, although Adams (1977) suggested that these are likely synonymous. Martins et al. (2016) recently revised this genus and outlined the confusion regarding the type locality of G. longicornis , which is stated as Georgia, but no geographic locality in South America matches that name; it is considered most likely a mislabelling, with the actual type locality being in northern South America.

Included species.

G. adspersus Navás, 1912 ( Brazil)

G. longicornis ( Walker, 1853) (likely South America)

Figures

FIGURE 2. Male genitalic morphology amongst different subfamilies of Osmylidae. Colour code: gonarcus (red), entoprocessus (blue), parameres (green), mediuncus (purple), hypandrium internum (orange). Scale line: 0.5 mm.

FIGURE 8. Gumillinae. Gumilla adspersus Navás, wings.

FIGURE 9. Gumillinae. Gumilla longicornis (Walker), wing venation (trichosors and vestiture omitted).