Trapelus sanguinolentus ( Pallas, 1814 )

Šmíd, Jiří, Moravec, Jiří, Kodym, Petr, Kratochvíl, Lukáš, Yousefkhani, Seyyed Saeed Hosseinian, Rastegar-Pouyani, Eskandar & Frynta, Daniel, 2014, Annotated checklist and distribution of the lizards of Iran, Zootaxa 3855 (1), pp. 1-97 : 13

Trapelus sanguinolentus ( Pallas, 1814)

TYPE. Not located ( Anderson 1999); Syntypes: ZMB 753-758 , ZMB 54811-12 (T. s. aralensis) .

TYPE LOCALITY. Kum-Ankatar on Terek River , Russia .

DISTRIBUTION. S Russia, Dagestan, Central Asian Republics, N Iran.

DISTRIBUTION IN IRAN. Fig. 23 . So far known only from three localities in Golestan and one in Khorasan Razavi Prov. , but other animals from NE Iran determined by now as T. agilis are also likely to belong to this species .

HABITAT. Semideserts with scattered vegetation which the species uses as a refuge and to observe the surroundings.

REMARKS. Formerly treated as a subspecies of T. agilis , but the latest phylogenetic study indicates that these two are not even closely related (i.e. sister) taxa ( Pyron et al. 2013). The nominotypical subspecies occurs W of the Caspian Sea in isolation from the main distribution range in Central Asia, where the subspecies T. s. aralensis (Lichtenstein) occurs. Based on the distribution of both subspecies the Iranian populations should be assigned to the latter.

REFERENCES. Rastegar-Pouyani (1998a, 1999a, b, 2005); Ananjeva et al. (2006).

Figures

FIGURES 20–25. 20. Trapelus agilis. Blue square denotes the type locality for T. a. khuzistanensis, dashed line delimits approximate boundary between this and the nominotypical subspecies. 21. Trapelus persicus. 22. Trapelus ruderatus. 23. Trapelus sanguinolentus. 24. Anguis colchica. 25. Pseudopus apodus.