Malagiella andringitra, Ubick & Griswold, 2011

Ubick, Darrell & Griswold, Charles E., 2011, The Malagasy Goblin Spiders Of The New Genus Malagiella (Araneae, Oonopidae), Bulletin of the American Museum of Natural History 2011 (356), pp. 1-86 : 79

Malagiella andringitra , new species

Figures 268–300 , 472 , 482 , 487 ; maps 1–4; table 1

TYPE: Male holotype and female allotype sifted from leaf litter in rainforest, 38 km S Ambalavo, 22 ° 129S, 46 ° 589E, 1680 m, Res. Andringitra, Fianarantsoa Province, Madagascar (23 Oct 1993, B. Fisher), deposited in CAS (8 holotype: CASENT 9029653, PBI _OON 03234; ♀ allotype CASENT 9029669, PBI _OON 03376) .

ETYMOLOGY: The species is named after the type locality.

DIAGNOSIS: The male differs from others in the group by the large bulb-cymbium (about 0.7 patella-tibia length; but about 0.5 in other species) and the presence of a shallow dorsal groove proximally on the bulb (fig. 482). The female differs in having a smaller dorsal scute (fig. 279) and the epigynal region weakly sclerotized and the receptaculum short and relatively straight (W/L about 0.25) (figs. 296, 297).

MALE (PBI_OON 03234). Total length 1.53, carapace length 0.74, width 0.62. CEPHALOTHORAX: Carapace (figs. 269, 270, 274) as in M. ranomafana . Eyes as in M. ranomafana , but slightly smaller; eye area about 0.5 carapace width, length subequal to clypeus length (figs. 269, 273). Sternum (figs. 272, 472) as in M. ranomafana . ABDO- MEN (figs. 269, 271, 275–277) as in M. valterova , but with slightly smaller book lung covers. GENITALIA: Epigastric region with sperm pore small, rebordered (fig. 271). Palp with bulb-cymbium large, about 0.7 patellatibia length; bulb with dorsal depression (figs. 288–295, 482, map 3).

FEMALE (PBI_OON 03376). Total length 1.82 (1.80–1.82), carapace length 0.78, width 0.68 (0.66–0.68), N 5 2. CEPHALOTHO- RAX (figs. 278–280, 283, 284): Clypeus sloping forward in lateral view, shorter than eye row length (fig. 280). Eye area about 1.3 clypeus length (figs. 280, 283). ABDOMEN (figs. 281, 285–287): Pedicel tube short (fig. 286), scutum dorsal extension slightly less than pedicel diameter (fig. 286). Dorsal scutum smaller than carapace, covering about 0.6 abdomen length and width (fig. 279). Postepigastric scutum weakly sclerotized. LEGS: patella plus tibia I longer than carapace (fig. 280). GENITALIA: Epigynal region weakly sclerotized, receptaculum short and relatively straight, W/L about 0.25 (figs. 296–300, map 4).

OTHER MATERIAL EXAMINED: MADA- GASCAR: Fianarantsoa Province: Res. Andringitra, 38 km S Ambalavo, 22 ° 129S, 46 ° 589E, 1680 m, rainforest, sifting leaf litter, 23 Oct 1993 (B. Fisher, CASENT 9029654, PBI_OON 03375), 1♀.

DISTRIBUTION: Known only from the type locality in Res. Andringitra, Madagascar.

Figures

Figs. 268–277. Malagiella andringitra, new species, male (PBI_OON 03234). 268. Habitus, ventral view. 269. Same, dorsal view. 270. Same, lateral view. Scale bar 5 1 mm. 271. Abdomen, ventral view. 272. Cephalothorax, ventral view. 273. Same, anterior view. 274. Same, posterior view. 275. Abdomen, anterior view. 276. Same, lateral view. 277. Same, posterior view.

Figs. 278–287. Malagiella andringitra, new species, female (PBI_OON 03376). 278. Habitus, ventral view. 279. Same, dorsal view. 280. Same, lateral view. Scale bar 5 1 mm. 281. Abdomen, ventral view. 282. Cephalothorax, ventral view. 283. Same, anterior view. 284. Same, posterior view. 285. Abdomen, anterior view. 286. Same, lateral view. 287. Same, posterior view.

Figs. 288–295. Malagiella andringitra, new species, male (PBI_OON 03234). 288. Palp, femur to bulb, prolateral view. 289. Same, retrolateral view. 290. Same, dorsal view. 291. Same, ventral view. 292. Same, anterior view. 293. Same, posterior view. 294. Bulb, retrolateral view. 295. Palp, retrolateral view. Note: images are of the right palp and have been reversed to give a left perspective comparable to images of other species.

Figs. 296–300. Malagiella andringitra, new species, female (PBI_OON 03376). 296. Abdomen, ventral view. 297. Epigynal region, ventral view. 298. Same, cleared specimen, arrow to apodeme orifice. 299. Same, lateral view, focus on apodeme (arrow) and associated muscles (dashes). 300. Same, focus on receptaculum (arrow) showing orifice (dash).

Figs. 472–480. Malagiella species, sternum, ventrolateral view, males (472–477) and females (478–480). 472. M. andringitra, PBI_OON 03234. 473. M. valterova, PBI_OON 03461. 474. M. fisheri, PBI_OON 03452. 475. M. ranomafana, PBI_OON 01999. 476. M. vohiparara, PBI_OON 03078. 477. M. toliara, PBI_OON 03377. 478. M. ranomafana, PBI_OON 01999. 479. M. vohiparara, PBI_OON 03078. 480. M. toliara, PBI_OON 03221.

Figs. 481–486. Malagiella species, male palpus, femur to bulb, prolateral view. 481. M. ranomafana, PBI_OON 03224. 482. M. andringitra, PBI_OON 03234. 483. M. valterova, PBI_OON 03461. 484. M. vohiparara, PBI_OON 03078. 485. M. fisheri, PBI_OON 03452. 486. M. toliara, PBI_OON 03377. Scale bar (200 Mm) applies to all palpi.

Fig. 487. Size variation in Malagiella measured as a function of carapace length to carapace width. Plot is based on measurements of 26 male specimens (‘‘x’’) and 30 female specimens (‘‘o’’). Blue lines enclose male values for a given species and red lines female values; disjunct conspecific readings are connected by straight lines. Green dotted lines enclose the three species groups. Brown dashed lines refer to the size of female DS, with the large dashes (upper right) enclosing species with the largest scutes and small dashes (lower left) those with the smallest. Abbreviations for species names: amba 5 M.ambalavo, andr 5 M. andringitra, fish 5 M. fisheri, good 5 M. goodmani, niki 5 M. nikina, rana 5 M. ranavalona, rano 5 M. ranomafana, toli 5 M. toliara, valt 5 M. valterova, vohi 5 M. vohiparara.