Gabrius fuscolaterus, Motschulsky, 1860 a: 76
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publication ID |
https://doi.org/ 10.1206/0003-0090.265.1.5 |
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persistent identifier |
https://treatment.plazi.org/id/03C1F679-FF20-FEC9-FF4F-FF7B803FF927 |
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treatment provided by |
Felipe |
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scientific name |
Gabrius fuscolaterus |
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fuscolaterus Motschulsky, 1860a: 76 View in CoL ( Gabrius ; Type locality: montagnes de Noura-Ellia à Ceylan).
— Fauvel, 1903d: 271 ( Philonthus ; synonym of thermarum ).
— Fauvel, 1905e: 83 ( Philonthus ; synonym of thermarum ).
— Bernhauer and Schubert, 1914: 357 ( Philonthus ; synonym of thermarum ).
— Cameron, 1932: 134 ( Philonthus ; synonym of thermarum ).
— Blackwelder, 1943: 403 ( Philonthus ; synonym of thermarum ).
— Scheerpeltz, 1957c: 234 ( Philonthus ; synonym of thermarum ).
— Smetana, 1958: 217 ( Philonthus ; synonym of thermarum ).
— Coiffait, 1974: 114 ( Gabronthus ; synonym of thermarum ).
— Scheerpeltz, 1978: 224 ( Philonthus ; synonym of thermarum ).
— Smetana, 1995e: 481 ( Gabronthus ; synonym of thermarum ).
turneri Tottenham, 1949b: 344 ( Philonthus ; Type locality: Natal: Van Reengen, Drakensberg). — Tottenham, 1955: 188 ( Gabronthus ; characters).
- DISTRIBUTION: South Africa.
ziloanus Levasseur, 1964: 292 ( Gabronthus ; Type locality: env. de Kolwezi, Katanga, Zilo).
- DISTRIBUTION: Congo.
[12 species; Ethiopian Region]
Glyphesthus Kraatz, 1858: 364 (species included: rufipennis , niger ). TYPE SPECIES: Glyphesthus rufipennis Kraatz , fixed by subsequent designation by Blackwelder, 1952: 171.
— Gemminger and Harold, 1868: 573 (catalog; cited as Glyphestes ).
— Bernhauer and Schubert, 1916: 411 (world catalog; 2 species).
— Bernhauer, 1931b: 85 (key to species).
— Scheerpeltz, 1933: 1421 (world catalog supplement).
Aleioglyphesthus Herman, 2001: 8 (subgenus of Glyphesthus ; species included: see Scheerpeltz, 1975). TYPE SPECIES: Glyphesthus (Aleioglyphesthus) zimmeri Scheerpeltz , fixed by original designation).
— Scheerpeltz, 1975: 110 (subgenus of Glyphesthus ; species included: congoensis , neavei , hauseri , zimmeri ; [Note: No type species designation so name was unavailable (Article 13.3)]).
congoensis Bernhauer, 1931b: 85 , 87 ( Glyphesthus ; Type locality: Belgisch-Kongo: Elisabethville).
— Scheerpeltz, 1933: 1421 ( Glyphesthus ; catalog).
— Scheerpeltz, 1975: 111 ( Glyphesthus ; subgenus Aleioglyphesthus ; characters; Congo).
- DISTRIBUTION: Congo].
evansi Bernhauer, 1931b: 85 , 86 ( Glyphesthus ; Type locality: Belgisches Kongogebiet:
18 m. S. W von Elisabethville).
— Scheerpeltz, 1933: 1421 ( Glyphesthus ; catalog).
— Scheerpeltz, 1975: 109 ( Glyphesthus ; characters; Congo).
- DISTRIBUTION: Congo.
hauseri Bernhauer, 1937: 305 ( Glyphesthus , cited as Glyphestus ; Type locality: Tanganyika: Uheheland, Kidugala).
— Scheerpeltz, 1975: 111 ( Glyphesthus ; subgenus Aleioglyphesthus ; characters; Kenya; Urundi).
- DISTRIBUTION: Kenya, Tanzania, Burundi.
leakeyi Bernhauer, 1937: 305 ( Glyphesthus ; cited as Glyphestus ; Type locality: Kenya: Oldoway, Turrei).
— Scheerpeltz, 1975: 110 ( Glyphesthus ; characters; Kenya).
- DISTRIBUTION: Kenya.
modestus Bernhauer, 1929f: 123 ( Glyphesthus ; Type locality: Haut-Uele: Tuku; Moto).
— Bernhauer, 1931b: 85 ( Glyphesthus ; characters in key; Belgian Congo).
— Scheerpeltz, 1933: 1421 ( Glyphesthus ; catalog).
— Scheerpeltz, 1975: 108 ( Glyphesthus ; characters; Congo region).
neavei Bernhauer, 1927: 57 ( Glyphesthus ; Type locality: Nyassaland: Mlanje ).
— Bernhauer, 1931b: 85 ( Glyphesthus ; characters in key; Nyassaland).
— Scheerpeltz, 1933: 1421 ( Glyphesthus ; catalog).
— Scheerpeltz, 1975: 111 ( Glyphesthus ; subgenus Aleioglyphesthus ; characters; Kenya).
- DISTRIBUTION: Kenya, Malawi.
niger Kraatz, 1858: 365 ( Glyphesthus ; Type locality: Senegambia).
— Fauvel, 1899a: 32 ( Glyphesthus ; Senegambia).
— Bernhauer and Schubert, 1916: 411 ( Glyphesthus ; catalog).
— Bernhauer, 1931b: 85 ( Glyphesthus ; characters in key; Senegambien).
— Scheerpeltz, 1933: 1421 ( Glyphesthus ; catalog; Ethiopia).
— Scheerpeltz, 1975: 110 ( Glyphesthus ; characters; Guinea; Senegal; Niger; Ivory Coast).
— Levasseur, 1968a: 54 ( Glyphesthus ; Central African Republic).
- DISTRIBUTION: “Senegambia”, Guinea, Ivory Coast, Niger, Central African Republic, Ethiopia.
nigerrimus Cameron, 1951c: 407 ( Glyphesthus ; Type locality: Kenya Colony : Kitui).
- DISTRIBUTION: Kenya.
parcepunctatus Bernhauer, 1927 g: 229 ( Glyphesthus ; Type locality: Harrar, Abessynien).
— Bernhauer, 1931b: 85 ( Glyphesthus ; characters in key; Abessinien).
— Scheerpeltz, 1933: 1421 ( Glyphesthus ; catalog).
— Scheerpeltz, 1975: 107 ( Glyphesthus ; characters; Abyssinia).
- DISTRIBUTION: Ethiopia.
picipennis Roth, 1851: 118 ( Acylophorus ; Type locality: Abyssinien).
— Fauvel, 1899a: 32 ( Glyphesthus ; Senegambie portugaise; Abyssinia; Tigre).
— Bernhauer and Schubert, 1916: 411 ( Glyphesthus ; catalog).
— Bernhauer, 1931b: 85 ( Glyphesthus ; characters in key; Abessinia).
— Scheerpeltz, 1933: 1421 ( Glyphesthus ; catalog).
— Scheerpeltz, 1975: 109 ( Glyphesthus ; characters; Guinea; Senegal; Niger; Cameroon; Aequatoria; Abyssinia).
- DISTRIBUTION: “Senegambia”, Senegal, Guinea, Niger, Cameroon, “Equatoria”, Ethiopia.
rufipennis Kraatz, 1858: 365 ( Glyphesthus ; Type locality: Senegambia).
— Bernhauer and Schubert, 1916: 411 ( Glyphesthus ; synonym of picipennis ).
rufipennis Kraatz, 1858 , see: picipennis Roth, 1851 .
sanguinipennis Bernhauer, 1931b: 85 , 86 ( Glyphesthus ; Type locality: Mashonaland; Natal: Colenso; Delagoabai ).
— Scheerpeltz, 1933: 1421 ( Glyphesthus ; catalog).
— Scheerpeltz, 1975: 108 ( Glyphesthus ; characters; South Africa).
- DISTRIBUTION: South Africa, Zimbabwe.
zimmeri Scheerpeltz, 1975: 103 , 112 ( Glyphesthus ; subgenus Aleioglyphesthus ; Type locality: Umgebung von Ugano im Matengo-Hochlande, West-Kenyas).
— Herman, 2001: 8 ( Glyphesthus ; subgenus Aleioglyphesthus ; type species of subgenus).
- DISTRIBUTION: Kenya.
[1 species; Oriental Region]
Hesperomimus Cameron, 1937: 17 (species included: abdominalis ; [Note: For the generic description Cameron spelled the name Hesperominus and for the species, Hesperomimus . Blackwelder, 1952: 183, argued that Hesperomimus is the correct spelling]). TYPE SPECIES: Hesperomimus abdominalis Cameron , fixed by monotypy.
— Blackwelder, 1952: 183 (type species: abdominalis ).
abdominalis Cameron, 1937: 17 ( Hesperomimus ; Type locality: Java: Batoerraden ,
G. Slamat).
[3 species; Palaearctic Region]
Hesperopalpus Shibata, 1973: 23 (species included: venustus ). TYPE SPECIES: Hesperopalpus venustus Shibata , fixed by original designation and monotypy.
— Shibata, 1986a: 124 (supplemental list of species from Taiwan).
— Schillhammer, 1999c: 303 (synopsis of genus; additional characters; notes; key to species). hesperiformis Cameron, 1932: 153 ( Philonthus ; Type locality: Nilgiri Hills).
— Scheerpeltz, 1933: 1345 ( Philonthus ; catalog).
— Schillhammer, 1996a: 83 ( Hesperopalpus ; characters; India).
— Schillhammer, 1999c: 307 ( Hesperopalpus ; characters; India).
- DISTRIBUTION: India.
venustus Shibata, 1973: 25 ( Hesperopalpus ; Type locality: Taiwan: Koantauchi , Nantou Hsien).
— Schillhammer, 1999c: 304 ( Hesperopalpus ; characters; Taiwan).
- DISTRIBUTION: Taiwan.
vietnamensis Schillhammer, 1999c: 306 ( Hesperopalpus ; Type locality: Vietnam: Taj Nguen, Bud Loij, 65 km N. Ankche).
- DISTRIBUTION: Vietnam.
[201 species; Neotropical, Nearctic, Palaearctic, Ethiopian,
Oriental, Australian, and Oceanic Regions]
Hesperus Fauvel, 1874: 200 [= 1874a: 426] (species included: rufipennis , baltimorensis , apicialis [cited as apicalis ]). TYPE SPECIES: Staphylinus rufipennis Gravenhorst , fixed by subsequent designation by R. Lucas, 1920: 324.
— Fauvel, 1875a: XXIX [= 1875b: 231] (catalog).
— Mulsant and Rey, 1876: 559 [= 1877: 415] (characters).
— Duvivier, 1883: 150 (catalog).
— Olliff, 1887: 508 (characters; Australian species).
— Heyden, Reitter, and Weise, 1891: 102 (list of species of Europe and Caucasus).
— Ganglbauer, 1895: 461 (characters).
— Fauvel, 1903d: 269 (key to New Caledonia species).
— Heyden, Reitter, and Weise, 1906: 163 (list of species of Europe and Caucasus).
— Reitter, 1909: 116, 122 (characters).
— Petri, 1912: 71 (list of species of Siebenbürgen).
— Bernhauer and Schubert, 1914: 363 (world catalog; 25 species).
— Leng, 1920: 108 (catalog of North American species).
— Cameron, 1921a: 354, 405 (characters; key to Singapore species; catalog).
— Gridelli, 1924 (characters; anatomy; revision of species; key to species).
— Winkler, 1925: 381 (catalog for Palaearctic region).
— Cameron, 1925: 64 (catalog of Indian species).
— Porta, 1926: 100 (characters; species of Italy).
— Bernhauer, 1928g: 116 (key to African species).
— Bruch, 1928a: 428 (catalog; Argentina).
— Roubal, 1930: 378 (catalog; Slovakia).
— Cameron, 1932: 156 (characters; key to species of British India).
— Scheerpeltz, 1933: 1371 (world catalog supplement).
— Scheerpeltz, 1936a: 488 (key to Nearctic and Neotropical species).
— Blackwelder, 1944: 138 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 184 (type species).
— Scheerpeltz, 1956b: 21 (key to African species).
— Moore, 1958 (characters; notes; key to North American species).
— Smetana, 1958: 258 (characters; habitus of one species; type species: rufipennis ; Czechoslovakian species). — Coiffait, 1963: 8, 18 (characters). — Székessy, 1963: 56 (characters). — Lohse, 1964: 174 (characters; key to central European species). — Moore, 1965: 98, 101 (characters; notes). — Scheerpeltz, 1968a: 79 (catalog of Austrian species). — Scheerpeltz, 1971 (key to species). — Last, 1973 (characters). — Tikhomirova, 1973a: 185 (checklist of species of USSR). — Coiffait, 1974: 320 (characters; species of western Palaearctic region). — Moore and Legner, 1975: 16 (catalog; North America north of Mexico). — Moore and Legner, 1979: 22 (characters; notes). — Burakowski, Mroczkowski, and Stefańska, 1980: 33 (catalog; Poland). — Shibata, 1983: 137 (checklist of species of Japan). — Hammond, 1984: 205 (checklist of species of Borneo). — Tóth, 1984: 41 (characters; habitus; species of Hungary). — Shibata, 1986a: 123 (supplemental list of species from Taiwan). — Shibata, 1990: 212 (key to species of Taiwan). — Smetana, 1995e: 32, 486 (characters; notes; key to North American species; phylogenetic relationships). — Ciceroni and Zanetti, 1995: 27 (list of species of Italy). — Downie and Arnett, 1996: 383 (characters; key to species of Northeastern North America). — Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species). — Newton, Thayer, Ashe, and Chandler, 2000: 396 (6 Nearctic species; notes; characters in key). — Hayashi, 2000: 23 (notes on empodial setae).
Hesperotropis Gridelli , 1924: 196 (subgenus of Hesperus ; species included: perfoliatus ; [Note: Following the description of the subgenus Hesperotropis Gridelli described Hesperus perfoliatus and that species was followed by Hesperus borneensis , H. bakeri , H. modestus , H. moultoni , H. ornatus , H. phaenomenalis , H. semicoeruleus , and H. terminalis but it is unclear that Gridelli intended them to be assigned to the new subgenus. In his key to species (page 182) Gridelli did not group H. perfoliatus in any special way with the other species and Scheerpeltz, 1933: 1371 and 1971: 193 included only H. perfoliatus in the subgenus]). TYPE SPECIES: Hesperus (Hesperotropis) perfoliatus Gridelli , fixed by monotypy [Note: If only one species was originally included then the type species is fixed by monotypy; if the other eight species were included then Blackwelder, 1952: 184, fixed the type species by subsequent designation]. — Cameron, 1932: 156 (subgenus of Hesperus ). — Scheerpeltz, 1933: 1371 (subgenus of Hesperus ). — Blackwelder, 1952: 184 (type species). — Scheerpeltz, 1971: 193 (subgenus of Hesperus ; characters).
abnormalis Cameron, 1952: 252 ( Hesperus ; Type locality: New Guinea: Snow Mountains: Moss Forest Camp, 2800 m). — Last, 1987: 49 ( Hesperus ; Papua New Guinea). - DISTRIBUTION: New Guinea.
addisadebaensis Scheerpeltz, 1971: 184 ( Hesperus ; Type locality: Aethiopien: Addisadeba). - DISTRIBUTION: Ethiopia.
aeneipennis Cameron, 1950c: 95 ( Hesperus ; Type locality: Belgian Congo: Mayidi). — Scheerpeltz, 1971: 188 ( Hesperus ; characters). - DISTRIBUTION: Congo.
aeneus Gridelli, 1924: 183 , 191 ( Hesperus ; Type locality: Sumatra occid., Si Rambé). — Scheerpeltz, 1933: 1371 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 166 ( Hesperus ; characters). - DISTRIBUTION: Indonesia.
africanus Bernhauer, 1915d: 145 ( Hesperus ; Type locality: Erythraea: Ghinda).
— Bernhauer, 1928g: 117 ( Hesperus ; characters in key).
— Scheerpeltz, 1933: 1371 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 24 ( Hesperus ; characters; Erythrea).
— Scheerpeltz, 1971: 185 ( Hesperus ; characters).
— Scheerpeltz, 1974c: 24 ( Hesperus ; Sudan).
- DISTRIBUTION: Eritrea, Sudan.
alboguttatus Cameron, 1946: 11 ( Hesperus ; Type locality: Philippines: Mindanao : Surigao).
— Scheerpeltz, 1971: 161 ( Hesperus ; characters).
- DISTRIBUTION: Philippines.
amabilis Kraatz, 1859: 97 ( Philonthus ; Type locality: India borealis).
— Bernhauer and Schubert, 1914: 328 ( Philonthus ; catalog).
— Cameron, 1932: 152 ( Philonthus ; characters; India).
— Scheerpeltz, 1933: 1332 ( Philonthus ; catalog).
— Schillhammer, 1999a: 62 ( Hesperus ).
- DISTRIBUTION: India.
amans Last, 1987: 33 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE: Wau, 1700 m).
- DISTRIBUTION: New Guinea.
anchora Fauvel, 1877: 258 ( Cafius ; Type locality: Nouvelle Calédonie).
— Fauvel, 1889b: 263 ( Hesperus ; New Caledonia).
— Fauvel, 1903d: 269 ( Hesperus ; New Caledonia).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 178 ( Hesperus ; characters).
- DISTRIBUTION: New Caledonia.
andrewesi Cameron, 1932: 157 ( Hesperus ; Type locality: Nilgiri Hills).
— Scheerpeltz, 1933: 1371 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 167 ( Hesperus ; characters).
- DISTRIBUTION: India.
apicialis Say, 1830: 37 [= 1834: 451] ( Staphylinus ; Type locality: Middle States [Note: See neotype designation by Smetana , 1995e]. Originally cited from the United States).
— Erichson, 1840: 931 ( Philonthus ; cited as apicalis ; [Note: Bernhauer and Schubert, 1914: 364 cited apicalis Erichson as a synonym of apicialis ; Erichson did not describe the species as new, he attributed it to Say but misspelled it]).
— Fauvel, 1874: 200 [= 1874a: 426] ( Hesperus ; cited as apicalis ).
— Horn, 1884: 220 ( Philonthus ; characters; Canada to Georgia).
— Blatchley, 1910: 390 ( Philonthus ; characters; Indiana).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Scheerpeltz, 1936a: 489 ( Hesperus ; characters in key).
— Moore, 1958: 315 ( Hesperus ; characters; Ohio, Missouri, Alabama).
— Scheerpeltz, 1971: 189 ( Hesperus ; characters).
— Frank, 1984: 478 ( Hesperus ; characters; Florida).
— Campbell and Davies, 1991: 119 ( Hesperus ; checklist of species; Canada).
— Smetana, 1995e: 498 ( Hesperus ; neotype designation: Middle States ; characters; notes; Ontario; Quebec; Alabama; Connecticut; Illinois; Indiana; Iowa; Kansas; Kentucky; Maine; Maryland; Massachusetts; Michigan; Nebraska; New Hampshire; New Jersey; New York; North Carolina; Ohio; Pennsylvania; Rhode Island; Texas; Virginia; West Virginia) .
— Downie and Arnett, 1996: 383 ( Hesperus ; characters; USA).
- DISTRIBUTION: Canada, USA.
haematurus Erichson, 1840: 504 ( Philonthus ; Type locality: America septentrionalis; [Note: See lectotype designation by Smetana, 1995e]).
— Erichson, 1840: 931 ( Philonthus ; synonym of apicialis ).
— LeConte, 1859: 566 ( Philonthus ; synonym of apicialis ).
— Frank, 1984: 478 ( Hesperus ; characters; Florida).
— Smetana, 1995e: 492 ( Hesperus ; lectotype designation; characters; notes; Alabama; Arkansas; Connecticut; Delaware; Georgia; Illinois; Indiana; Iowa; Kansas; Kentucky; Louisiana; Maryland; Massachusetts; Michigan; Mississippi; Missouri; Montana; Nebraska; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; South Carolina; Virginia) .
— Downie and Arnett, 1996: 383 ( Hesperus ; characters; USA).
- DISTRIBUTION: Canada, USA.
barrioni Last, 1991: 1076 ( Hesperus ; Type locality: Philippines: Luzon Island: Siniloan, Laguna, Magsaysay village ).
- DISTRIBUTION: Philippines.
belonuchiformis Bernhauer, 1915d: 145 ( Hesperus ; Type locality: Abessinien: Boroda ).
— Bernhauer, 1928g: 117 ( Hesperus ; characters in key).
— Scheerpeltz, 1933: 1371 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 23 ( Hesperus ; characters; Abyssinia).
— Scheerpeltz, 1971: 184 ( Hesperus ; characters).
- DISTRIBUTION: Ethiopia.
bicolor Smetana, 1995e: 497 ( Hesperus ; Type locality: Florida: Coronado Bch.).
- DISTRIBUTION: USA.
bifuscipennis Lea, 1931: 373 ( Hesperus ; Type locality: Papua: Mount Lamington ).
— Scheerpeltz, 1933: 1371 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 177 ( Hesperus ; characters).
- DISTRIBUTION: New Guinea.
birmanus Cameron, 1932: 160 ( Hesperus ; Type locality: Burma: Karen Mts.; Assam: Sadiya). — Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 167 ( Hesperus ; characters).
- DISTRIBUTION: India, Myanmar.
borneensis Bernhauer, 1915f: 142 ( Hesperus ; Type locality: Sarawak: Mt. Matang ,
2000 Fuss).
— Gridelli, 1924: 198 ( Hesperus ; characters; Sarawak).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 167 ( Hesperus ; characters).
- DISTRIBUTION: Borneo, Indonesia.
rufipennis Cameron, 1918b: 171 ( Leucitus ; [preoccupied]; Type locality: Borneo, Quop, W. Sarawak; also unnamed variety from Bedagei , Sumatra).
— Scheerpeltz, 1933: 1372 ( Hesperus ; synonym of borneensis ).
bruchi Bernhauer, 1921c: 169 ( Hesperus ; Type locality: Argentinien: Prov. Santiago d. Estero). — Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1936a: 490 ( Hesperus ; characters in key).
— Scheerpeltz, 1971: 191 ( Hesperus ; characters).
- DISTRIBUTION: Argentina.
brunneipennis Scheerpeltz, 1971 [Note: Nomen nudum], see: rufipennis Gravenhorst, 1802 . buluensis Last, 1973: 432 ( Hesperus ; Type locality: New Guinea: Garaina, Bulu Valley).
- DISTRIBUTION: New Guinea.
nexilis Last, 1973: 432 ( Hesperus ; variety of buluensis ; [Note: Nomen nudum (Article 15.2)]; cited from New Guinea: Upper Wario, 5000 ft.).
burgeoni Bernhauer, 1928 g: 119 ( Hesperus ; Type locality: Belgisch-Kongo: Haut-Uele: Moto; Watsa).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 26 ( Hesperus ; characters; Belgian Congo).
— Scheerpeltz, 1971: 183 ( Hesperus ; characters).
cafioides Fauvel, 1889c: 187 ( Hesperus ; Type locality: Liberia Africae ).
— Fauvel, 1903e: 240 ( Hesperus ; Cameroon; Liberia; Assinie; Angola; Gabon; Congo; Sierra Leone).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Bernhauer, 1928g: 117 ( Hesperus ; characters in key).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Bernhauer, 1942c: 363 ( Hesperus ; Cameroon).
— Scheerpeltz, 1956b: 23 ( Hesperus ; characters; Liberia; Nigeria; Togo; Cameroon; French Congo; Belgian Congo).
— Scheerpeltz, 1971: 187 ( Hesperus ; characters).
— Scheerpeltz, 1971c: 170 ( Hesperus ; Cameroon).
- DISTRIBUTION: Guinea, Sierra Leone, Liberia, Ivory Coast, Togo, Nigeria, Cameroon, Gabon, French Equatorial Africa, Congo, Angola.
laniger Fauvel, 1889b: 188 ( Hesperus ; Type locality: Stanley Pool , Congo).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Bernhauer, 1928g: 117 ( Hesperus ; variety of cafioides ; characters in key).
— Scheerpeltz, 1933: 1372 ( Hesperus ; variety of cafioides ).
— Scheerpeltz, 1956b: 23 ( Hesperus ; cited as valid species; characters; Belgian Congo).
— Levasseur, 1966a: 1214 ( Hesperus ; Cameroon; Guinea; Ivory Coast; Congo).
— Scheerpeltz, 1971: 187 ( Hesperus ; variety of cafioides ; characters).
candidus Last, 1968: 353 ( Philonthus ; Type locality: New Guinea: Okapa).
— Last, 1973: 425 ( Hesperus ).
— Last, 1987: 49 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
caulis Last, 1981: 128 ( Hesperus ; Type locality: Goroka).
— Last, 1987: 49 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
chinensis Cameron, 1940d: 252 ( Hesperus ; Type locality: China).
- DISTRIBUTION: China.
cingulatus Cameron, 1926 , see: feae Fauvel, 1895 .
coelestinus Bernhauer, 1926b: 41 ( Hesperus ; Type locality: Luzon: Balbalan).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 174 ( Hesperus ; cited as coelestis ; characters).
- DISTRIBUTION: Philippines.
colmani Last, 1987: 33 ( Hesperus ; Type locality: Papua Guinea: New Guinea NE:
Mount Missim, 2100 m).
- DISTRIBUTION: New Guinea.
coloratus Last, 1987: 34 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE: Torricelli Mts., Mobetei, 750 m).
- DISTRIBUTION: New Guinea.
conciliatus Last, 1981: 129 ( Hesperus ; Type locality: Daulo Pass, Asato-Chimbu Divide).
- DISTRIBUTION: New Guinea.
confusus Last, 1981: 128 ( Hesperus ; Type locality: Goroka, 8400 feet).
- DISTRIBUTION: New Guinea.
copiosus Last, 1968: 359 ( Philonthus ; Type locality: New Guinea: Okapa).
— Last, 1973: 425 ( Hesperus ).
— Last, 1987: 49 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
delectus Last, 1968: 365 ( Philonthus ; Type locality: New Guinea: Purosa).
— Last, 1973: 425 ( Hesperus ; characters).
- DISTRIBUTION: New Guinea.
depressus Bernhauer, 1908c: 109 ( Hesperus ; Type locality: Kamerun).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Bernhauer, 1928g: 119 ( Hesperus ; characters in key; Belgian Congo; Cameroon).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 182 ( Hesperus ; characters).
- DISTRIBUTION: Cameroon, Congo.
diabolicus Bernhauer, 1926b: 42 ( Hesperus ; Type locality: Luzon: Mt. Polis).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 170 ( Hesperus ; characters).
- DISTRIBUTION: Philippines.
distincticornis Cameron, 1937 , see: roepkei Bernhauer, 1911 .
divinus Bernhauer, 1915 g: 190 ( Hesperus ; Type locality: Deutsch-Neuguinea: Lordberg).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 180 ( Hesperus ; characters).
- DISTRIBUTION: New Guinea.
dohertyi Cameron, 1942c: 845 ( Hesperus ; Type locality: S. Celebes: Tamba ).
- DISTRIBUTION: Indonesia.
drescheri Cameron, 1937: 16 ( Hesperus ; Type locality: Java: Batoerraden, G. Slamat; G. Tangkoeban Prahoe, alt. 4000–5000 ft).
— Scheerpeltz, 1971: 165 ( Hesperus ; characters).
- DISTRIBUTION: Indonesia.
effectus Last, 1987: 34 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE: Wau, 1200 m).
- DISTRIBUTION: New Guinea.
eidmanni Scheerpeltz, 1936a: 484 , 491 ( Hesperus ; Type locality: Brasilien: Umgebung von Mendes, E. d. Rio ).
— Scheerpeltz, 1971: 193 ( Hesperus ; characters).
- DISTRIBUTION: Brazil.
excellens Cameron, 1952: 250 ( Hesperus ; Type locality: Misool Island: Fakal).
- DISTRIBUTION: Indonesia.
excelsus Cameron, 1931 , see: Philonthus .
feae Fauvel, 1895: 260 ( Hesperus ; Type locality: Bírmanie, Carin Cheba , 900–1100 m;
Carin Asciuii Ghécu, 1400–1500 m; Sikkim, Darjeeling).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Gridelli, 1924: 189 ( Hesperus ; characters; Burma; India).
— Cameron, 1932: 158 ( Hesperus ; characters; Burma; India).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 155, 157 ( Hesperus ; characters).
— Schillhammer, 1999a: 62 ( Hesperus ; species may be synonymous with amabilis Kraatz but further study is needed).
- DISTRIBUTION: Myanmar, India.
cingulatus Cameron, 1926b: 361 ( Hesperus ; Type locality: Mussorie District, Arni Gad. Chakrata District, Bangar. Simla Hills, Theog, 6000 to 7600 feet).
— Cameron, 1932: 158 ( Hesperus ; synonym of feae ).
— Scheerpeltz, 1933: 1372 ( Hesperus ; variety of feae ).
— Scheerpeltz, 1971: 155, 158 ( Hesperus ; aberration of feae ; characters).
ferrugineus Bernhauer, 1925: 36 ( Hesperus ; Type locality: Argentinien: Alta Gracia
La Granja, Sierra de Cordoba).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1936a: 490 ( Hesperus ; characters in key).
— Scheerpeltz, 1971: 192 ( Hesperus ; characters).
flavoris Last, 1987: 36 ( Hesperus ; Type locality: Irian: Vogelkop, Fak Fak, S. coast of Bomberai, 100–700 m).
- DISTRIBUTION: New Guinea.
fraternus Bernhauer, 1928 g: 117 ( Hesperus ; Type locality: Natal: Frere; Zululand).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 24 ( Hesperus ; characters; Natal; Zululand).
— Scheerpeltz, 1971: 185 ( Hesperus ; characters).
- DISTRIBUTION: South Africa.
fraudator Last, 1981: 133 ( Hesperus ; Type locality: Madang District, Finisterre Mts., Budemu , c. 4000 feet).
- DISTRIBUTION: New Guinea.
gagativentris Lea, 1931: 374 ( Hesperus ; Type locality: Papua: Mount Lamington ).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 179 ( Hesperus ; characters).
- DISTRIBUTION: New Guinea.
gestroi Bernhauer, 1915 k: 302 ( Hesperus ; Type locality: Insel Fernando Poo: Moka , 1300–1500 m).
— Bernhauer, 1915o: 363 ( Hesperus ; Fernando Poo).
— Bernhauer, 1928g: 116 ( Hesperus ; characters in key).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 21 ( Hesperus ; characters).
— Scheerpeltz, 1971: 189 ( Hesperus ; characters).
- DISTRIBUTION: Equatorial Guinea.
giluensis Last, 1987: 36 ( Hesperus ; Type locality: Papua New Guinea: Mount Giluwe , 2800–3200 m).
- DISTRIBUTION: New Guinea.
glaucus Last, 1981: 133 ( Hesperus ; Type locality: New Guinea: Lufa).
— Last, 1987: 49 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
gloriosus Lea, 1925: 227 ( Hesperus ; Type locality: Queensland: Coen River; Cairns District). — Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 179 ( Hesperus ; characters).
- DISTRIBUTION: Australia.
graciliventris Cameron, 1952: 254 ( Hesperus ; Type locality: New Guinea: Snow Mountains: Araucaria Camp, 800 m).
— Last, 1987: 48 ( Hesperus ; Papua New Guinea; Irian).
- DISTRIBUTION: New Guinea.
obscuritus Last, 1987: 48 ( Hesperus ; variety of graciliventris ; [Note: Nomen nudum (Article 15.2)]; cited from Papua New Guinea: Wau, Big Wau Creek, 1300 m; Morobe District, Wau, 1300 m; Anga Gorge E of Mendi; Huon Pen., Pindiu. Irian: Vogelkop, bomberai, 700–900 m).
gratiosus Cameron, 1943a: 344 ( Hesperus ; Type locality: New Guinea: Finsch Haven).
— Last, 1987: 49 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
gratus Cameron, 1943a: 342 ( Philonthus ; [preoccupied]; Type locality: New Guinea: Wareo, Finsch Haven).
— Last, 1987: 49 ( Hesperus ; Papua New Guinea).
— Herman, 2001: 45 ( Hesperus ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
- DISTRIBUTION: New Guinea.
gressitti Last, 1987: 36 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE:
Karimui, south of Goroka, 1000 m).
gridellianus Bernhauer, 1928 g: 117 ( Hesperus ; Type locality: Portugiesisch-Guinea:
Rio Cassine; Bahia de San Carlos ).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 22 ( Hesperus ; characters).
— Scheerpeltz, 1971: 188 ( Hesperus ; characters).
- DISTRIBUTION: Guinea-Bissau.
gridellii Cameron, 1926 , see: Philonthus .
haematurus Erichson, 1840 , see: apicialis Say, 1830 .
haemorrhoidalis W. J. MacLeay, 1873: 140 ( Philonthus ; Type locality: Gayndah).
— Fauvel, 1877: 260 ( Hesperus ; characters; Australia).
— Olliff, 1887: 508 ( Hesperus ; characters; Australia).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Lea, 1925: 227 ( Hesperus ; characters; Australia).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 178 ( Hesperus ; characters).
- DISTRIBUTION: Australia.
mirabilis Fauvel, 1877: 260 ( Hesperus ; Type locality: Australie, Queensland, Wide Bay). — Fauvel, 1878g: 543 ( Hesperus ; Australia).
— Olliff, 1887: 508 ( Hesperus ; synonym of haemorrhoidalis ).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Bernhauer, 1920: 16 ( Hesperus ; Australia).
— Lea, 1925: 227 ( Hesperus ; synonym of haemorrhoidalis ).
— Scheerpeltz, 1933: 1372 ( Hesperus ; synonym of haemorrhoidalis ).
— Scheerpeltz, 1971: 178 ( Hesperus ; synonym of haemorrhoidalis ).
hagenensis Last, 1987: 37 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE: Tomba, slopes of Mt. Hagen, 2450 m).
- DISTRIBUTION: New Guinea.
heinrichi Last, 1975a: 6 ( Hesperus ; Type locality: Papua: Madang).
- DISTRIBUTION: New Guinea.
heynei Wendeler, 1927: 7 ( Hesperus ; Type locality: Costa Rica: Turrialba , 800–900 m).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1936a: 489 ( Hesperus ; characters in key).
— Scheerpeltz, 1971: 190 ( Hesperus ; characters).
- DISTRIBUTION: Costa Rica.
hilleri Last, 1987: 37 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE: Karimui, S of Goroka, 1000 m).
- DISTRIBUTION: New Guinea.
hirsutus Last, 1973: 431 ( Hesperus ; Type locality: New Guinea: Garaina).
— Last, 1987: 49 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
hobbyi Bernhauer, 1936e: 214 ( Hesperus ; Type locality: Sarawak: foot of Mt. Dulit , Junction of rivers Tinjar and Lejok).
— Scheerpeltz, 1971: 163 ( Hesperus ; characters).
- DISTRIBUTION: Malaysia.
holzschuhi Schillhammer, 1991: 52 ( Hesperus ; Type locality: E-Nepal: Koshi, Basantapur , 2300 m).
- DISTRIBUTION: Nepal.
honestus Last, 1981: 120 ( Philonthus ; Type locality: New Guinea: Mount Hagen).
— Last, 1987: 49 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
hornabrooki Last, 1968: 354 ( Philonthus ; Type locality: New Guinea: Okapa).
— Last, 1987: 49 ( Hesperus ; Papua New Guinea; Irian).
- DISTRIBUTION: New Guinea.
huonensis Last, 1981: 129 ( Hesperus ; Type locality: Tep Tep, Finnisterre Range, Huon Pen). - DISTRIBUTION: New Guinea.
inaequalis Fauvel, 1895: 258 ( Hesperus ; Type locality: Bírmanie, Carin Cheba , 900–1100 m; Carin Ghécu, 1300–1400 m).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Gridelli, 1924: 193 ( Hesperus ; characters; Burma).
— Cameron, 1932: 160 ( Hesperus ; characters; Burma).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1965: 108 ( Hesperus ; Burma).
— Scheerpeltz, 1971: 162 ( Hesperus ; characters).
- DISTRIBUTION: Myanmar.
indigaceus Fauvel, 1878 g: 544 ( Hesperus ; Type locality: Nouvelle-Calédonie, Ile des Pins). — Fauvel, 1889b: 263 ( Hesperus ; New Caledonia).
— Fauvel, 1903d: 269 ( Hesperus ; Iles des Pins).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1966e: 402 ( Hesperus ; characters; New Caledonia).
— Scheerpeltz, 1971: 181 ( Hesperus ; characters).
- DISTRIBUTION: New Caledonia.
ignoratus Ito, 1994c: 81 ( Hesperus ; Type locality: Japan: Kyoto Pref.: Ogoshi ).
- DISTRIBUTION: Japan.
interjectus Last, 1987: 37 ( Hesperus ; Type locality: Papua New Guinea NE: Karimui ,
S of Goroka, 1000 m. Also cited from Irian).
- DISTRIBUTION: New Guinea.
javanus Cameron, 1937: 15 ( Hesperus ; Type locality: Java: Batoerraden, G. Slamat).
— Scheerpeltz, 1971: 157 ( Hesperus ; characters).
- DISTRIBUTION: Indonesia.
junius Last, 1987: 38 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE:
Daulo Pass, 2500 m, Asaro-Chimbu Div.).
- DISTRIBUTION: New Guinea.
justus Last, 1968: 348 ( Philonthus ; Type locality: New Guinea: Okapa).
— Last, 1973: 425 ( Hesperus ).
— Last, 1987: 50 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
kaindiensis Last, 1980: 155 ( Hesperus ; Type locality: New Guinea: Wau , Mt. Kaindi, 2000 m). - DISTRIBUTION: New Guinea.
kokodanus Cameron, 1937a: 101 ( Philonthus ; Type locality: New Guinea: Kokoda,
alt. 1200 ft.).
— Last, 1987: 50 ( Hesperus ; cited as kakodanus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
kambaitiensis Scheerpeltz, 1965: 242 ( Hesperus ; Type locality: N.E. Burma, Kambaiti ,
7000 ft.).
— Scheerpeltz, 1971: 169 ( Hesperus ; characters).
- DISTRIBUTION: Myanmar.
kraatzi Eppelsheim, 1895a: 129 ( Hesperus ; Type locality: Togoland).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Bernhauer, 1928g: 116 ( Hesperus ; characters in key).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Scheerpeltz, 1956a: 485 ( Hesperus ; Guinea; Liberia; Nigeria; Cameroon; French Congo;
— Scheerpeltz, 1956b: 22 ( Hesperus ; characters).
— Scheerpeltz, 1971: 188 ( Hesperus ; characters).
- DISTRIBUTION: Guinea, Liberia, Nigeria, Cameroon, French Equatorial Africa, Congo, Uganda, Kenya, Tanzania.
laevigatus Fauvel, 1895: 259 ( Hesperus ; Type locality: Bírmanie, Carin Cheba, 900–1100 m; Pulo-Penang ; Malacca; Singapore; Sumatra ; Borneo; Java).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Cameron, 1921a: 378, 405 ( Hesperus ; characters; Singapore).
— Gridelli, 1924: 183 ( Hesperus ; characters; Burma; Sumatra).
— Bernhauer, 1928: 17 ( Hesperus ; Sumatra).
— Cameron, 1932: 157 ( Hesperus ; characters; Burma).
— Scheerpeltz, 1933: 1372 ( Hesperus ; catalog).
— Cameron, 1933d: 390 ( Hesperus ; Sumatra).
— Scheerpeltz, 1965: 108 ( Hesperus ; Burma).
— Scheerpeltz, 1971: 159 ( Hesperus ; characters).
— Frank, 1982: 36 ( Hesperus ; list of parasites).
- DISTRIBUTION: Myanmar, Malaysia, Singapore, Borneo, Indonesia.
sarawakensis Gridelli, 1924: 185 ( Hesperus ; variety of laevigatus ; Type locality: Borneo: Sarawak).
— Scheerpeltz, 1933: 1373 ( Hesperus ; variety of laevigatus ).
— Scheerpeltz, 1971: 159 ( Hesperus ; variety of laevigatus ; characters).
laniger Fauvel, 1889 , see: cafioides Fauvel, 1889 .
levitus Last, 1987: 38 ( Hesperus ; Type locality: Papua New Guinea NE: Morobe Distr., Wau, 1200 m. Also cited from New Britain).
- DISTRIBUTION: New Guinea, New Britain.
lifuanus Fauvel, 1903d: 269 ( Hesperus ; Type locality: Lifou).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 179 ( Hesperus ; characters).
- DISTRIBUTION: New Caledonia.
locus Last, 1987: 39 ( Hesperus ; Type locality: Papua New Guinea NE: Daulo Pass , Asaro-Chimbu Div., 2400 m).
- DISTRIBUTION: New Guinea.
longicornis Cameron, 1950e: 430 ( Hesperus ; Type locality: Cameroon: Ngauyanga ).
- DISTRIBUTION: Cameroon.
longipennis Cameron, 1952: 253 ( Hesperus ; Type locality: New Guinea: Snow Mountains: Lower Mist Camp, 1700 m).
- DISTRIBUTION: New Guinea.
luluanus Scheerpeltz, 1956b: 23 ( Hesperus ; Type locality: Belgisch-Kongo).
— Herman, 2001: 25 ( Hesperus ; synonymy).
- DISTRIBUTION: Congo.
luluanus Scheerpeltz, 1971: 184 ( Hesperus ; [preoccupied]; Type locality: Kongo Gebiet und Uganda: Kapanga).
— Herman, 2001: 25 ( Hesperus ; synonym of luluanus Scheerpeltz, 1956 ).
luluanus Scheerpeltz, 1971 , see: luluanus Scheerpeltz, 1956 .
luzonicus Bernhauer, 1926b: 43 ( Hesperus ; Type locality: Luzon: Balbalan).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 171 ( Hesperus ; characters).
- DISTRIBUTION: Philippines.
maai Last, 1987: 39 ( Hesperus ; Type locality: Irian: Vogelkop, Bomberai, 700–900 m).
- DISTRIBUTION: New Guinea.
— Scheerpeltz, 1971: 161 ( Hesperus ; characters).
- DISTRIBUTION: Philippines.
malaisei Scheerpeltz, 1965: 240 ( Hesperus ; Type locality: N.E. Burma, Kambaiti, 7000 ft.). — Scheerpeltz, 1971: 173 ( Hesperus ; characters).
- DISTRIBUTION: Myanmar.
malayanus Cameron, 1932a: 142 ( Hesperus ; Type locality: Selangor: Bukit Kutu ).
— Scheerpeltz, 1971: 159 ( Hesperus ; characters).
- DISTRIBUTION: Malaysia.
mandatus Last, 1987: 39 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE: Morobe Distr., Wau, 1200 m).
- DISTRIBUTION: New Guinea.
maris Last, 1987: 40 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE: Torricelli Mts., Mobitei, 750 m).
- DISTRIBUTION: New Guinea.
meyeri Bernhauer, 1928 g: 119 ( Hesperus ; Type locality: Uganda).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 26 ( Hesperus ; characters; Uganda).
— Scheerpeltz, 1971: 182 ( Hesperus ; characters).
- DISTRIBUTION: Uganda.
mirabilis Fauvel, 1877 , see: haemorrhoidalis W. J. MacLeay, 1873 .
mirificus Herman, 2001: 45 ( Hesperus ; replacement name for mirus Last ).
- DISTRIBUTION: New Guinea.
mirus Last, 1981: 132 ( Hesperus ; [preoccupied]; Type locality: Frigano, Okapa,
Eastern Highlands).
— Herman, 2001: 45 ( Hesperus ; synonym of mirificus ).
mirus Bernhauer, 1915d: 146 ( Hesperus ; Type locality: Kamerun).
— Bernhauer, 1915o: 363 ( Hesperus ; Cameroon).
— Bernhauer, 1928g: 116 ( Hesperus ; characters in key).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 21 ( Hesperus ; characters; Cameroon).
— Levasseur, 1968a: 52 ( Hesperus ; Central African Republic).
— Scheerpeltz, 1971: 187 ( Hesperus ; characters).
- DISTRIBUTION: Cameroon, Central African Republic.
mirus Last, 1981 , see: mirificus Herman, 2001 .
mixtus Last, 1968: 366 ( Philonthus ; Type locality: New Guinea: Karimu).
— Last, 1973: 425 ( Hesperus ).
— Last, 1981: 128 ( Hesperus ; New Guinea).
— Last, 1987: 50 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
modestus Bernhauer, 1915 h: 229 ( Hesperus ; Type locality: Java: Preanger ).
— Gridelli, 1924: 199 ( Hesperus ; characters; Java).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 170 ( Hesperus ; characters).
- DISTRIBUTION: Indonesia.
modulatus Last, 1968: 358 ( Philonthus ; Type locality: New Guinea: Buana , Sarawaged Range, 3000 ft., 20–30 miles north of Lae).
— Last, 1973: 425, 426 ( Hesperus ; characters).
— Last, 1987: 50 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
monstratus Last, 1987: 40 ( Hesperus ; Type locality: Papua New Guinea: Owen Stanley montanellus Bernhauer, 1926b: 43 ( Hesperus ; Type locality: Luzon: Mt. Banahao).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 171 ( Hesperus ; characters).
- DISTRIBUTION: Philippines.
moresbyensis Last, 1987: 42 ( Hesperus ; Type locality: Papua New Guinea: Bisianumu ,
E of Port Moresby, 500 m).
- DISTRIBUTION: New Guinea.
moroensis Last, 1981: 132 ( Hesperus ; Type locality: Madang district, Finisterre, Mts., Moro , c. 5550 feet).
- DISTRIBUTION: New Guinea.
moultoni Bernhauer, 1915f: 143 ( Hesperus ; Type locality: Britisch-Nord-Borneo:
Mt. Kinabalu).
— Gridelli, 1924: 199 ( Hesperus ; characters; Borneo).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 172 ( Hesperus ; characters).
- DISTRIBUTION: Borneo.
multipunctus Last, 1981: 130 ( Hesperus ; Type locality: New Hebrides: West Santo , Tartarii, 4000 ft.).
- DISTRIBUTION: Vanuatu.
natalensis Scheerpeltz, 1956b: 22 ( Hesperus ; Type locality: Natal).
— Herman, 2001: 25 ( Hesperus ; synonymy).
- DISTRIBUTION: South Africa.
natalensis Scheerpeltz, 1971: 188 ( Hesperus ; [preoccupied]; Type locality: Süd-Afrika: Natal).
— Herman, 2001: 25 ( Hesperus ; synonym of natalensis Scheerpeltz, 1956 ).
natalensis Scheerpeltz, 1971 , see: natalensis Scheerpeltz, 1956 .
nexilis Last, 1973 [Note: Nomen nudum], see: buluensis Last, 1973 .
nigriventris Gridelli, 1924: 182 , 186 ( Hesperus ; Type locality: Isola Nias , Sumatra).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 169 ( Hesperus ; characters).
- DISTRIBUTION: Indonesia.
nobilis Bernhauer, 1928 g: 119 ( Hesperus ; Type locality: Belgisch-Kongo: Bas, Uele).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 27 ( Hesperus ; characters; Belgian Congo).
— Scheerpeltz, 1971: 183 ( Hesperus ; characters).
- DISTRIBUTION: Congo.
novoteutonicus Wendeler, 1956a: 232 ( Hesperus ; Type locality: Brasilien: Nova Teutonia ).
- DISTRIBUTION: Brazil.
obesus Last, 1989: 878 ( Hesperus ; Type locality: Papua New Guinea: Onerunka , near Kainantu).
- DISTRIBUTION: New Guinea.
obscuricollis Cameron, 1941c: 379 ( Hesperus ; Type locality: Philippines: Mindanao : Surigao). - DISTRIBUTION: Philippines.
obscuricollis Scheerpeltz, 1971 , see: septuosus Herman, 2001 .
obscuripennis Scheerpeltz, 1971 [Note: Nomen nudum], see: bakeri Bernhauer, 1919 . obscurior Bernhauer, 1915d: 145 ( Hesperus ; variety of belonuchiformis ; Type locality: Abessinien: Errer; Djerer-Tal).
— Bernhauer, 1928g: 118 ( Hesperus ; characters in key).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 24 ( Hesperus ; characters; Abyssinia).
picticollis Gridelli, 1924: 183 , 194 ( Hesperus ; Type locality: Isole Mentawei; Sipor Sereinu). — Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 164 ( Hesperus ; characters).
- DISTRIBUTION: Indonesia.
picticornis Lea, 1925: 227 ( Hesperus ; Type locality: New South Wales: Tweed River).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 177 ( Hesperus ; characters).
- DISTRIBUTION: Australia.
plagiarius Last, 1987: 42 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE: Huon Peninsula, Pindiu ).
- DISTRIBUTION: New Guinea.
plumbeus Last, 1981: 127 ( Hesperus ; Type locality: Morobe Dist., Edie CK, Bulldog Road, 9700 ft.).
- DISTRIBUTION: New Guinea.
postulator Last, 1987: 42 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE: Morobe Distr., Wau, 1300 m).
- DISTRIBUTION: New Guinea.
preangeranus Cameron, 1937: 16 ( Hesperus ; Type locality: Java: G. Tangkoeban Prahoe,
alt. 4000–5000 ft).
— Scheerpeltz, 1971: 162 ( Hesperus ; characters).
- DISTRIBUTION: Indonesia.
primitus Last, 1987: 43 ( Hesperus ; Type locality: Irian: Waris S. of Jayapura, Hollandia, 450–500 m. Also cited from Papua New Guinea).
- DISTRIBUTION: New Guinea.
privus Last, 1973: 429 ( Hesperus ; Type locality: Okapa).
— Last, 1987: 50 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
probatus Last, 1987: 43 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE: Aiyura, 1600 m).
- DISTRIBUTION: New Guinea.
pseudobirmanus Scheerpeltz, 1965: 238 ( Hesperus ; Type locality: N.E. Burma, Kambaiti , 2000 m).
— Scheerpeltz, 1971: 168 ( Hesperus ; characters).
- DISTRIBUTION: Myanmar.
pseudoviolaceus Last, 1981: 130 ( Hesperus ; Type locality: Wantout, Huon Peninsular).
- DISTRIBUTION: New Guinea.
pulleinei Olliff, 1887: 512 ( Hesperus ; [Note: Olliff’s description was published before Blackburn’s]; Type locality: Burnside, Adelaide , South Australia).
— Blackburn, 1888: 7 ( Hesperus ; [Note: Bernhauer and Schubert, 1914: 364 cited Blackburn as the author for this name, and although Olliff also cited Blackburn as the author, Olliff’s description appeared first]; characters; Australia).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 180 ( Hesperus ; characters).
- DISTRIBUTION: Australia.
puncticeps Bernhauer, 1928 g: 118 ( Hesperus ; Type locality: Uganda: Entebbe ).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 25 ( Hesperus ; characters; Uganda).
— Scheerpeltz, 1971: 186 ( Hesperus ; characters).
- DISTRIBUTION: Uganda.
punctipennis Cameron, 1941c: 380 ( Hesperus ; Type locality: Philippines: Tangcolan :
Bukidnon).
purpuripennis Bernhauer, 1915 g: 189 ( Hesperus ; Type locality: Ralums).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 180 ( Hesperus ; characters).
— Last, 1987: 50 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
quinquecolor Lea, 1931: 372 ( Hesperus ; Type locality: New Guinea: Wareo).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 180 ( Hesperus ; characters).
- DISTRIBUTION: New Guinea.
rarus Last, 1989: 880 ( Hesperus ; Type locality: Papua New Guinea: Kainantu , Onerunka).
- DISTRIBUTION: New Guinea.
raynori Last, 1987: 44 ( Hesperus ; Type locality: Papua New Guinea: Owen Stanley Range , Goilala, Loloipa).
- DISTRIBUTION: New Guinea.
ricardi Last, 1973: 431 ( Hesperus ; Type locality: New Guinea: Upper Wario, 5000 ft.).
— Last, 1987: 50 ( Hesperus ; cited as richardi; Papua New Guinea).
- DISTRIBUTION: New Guinea.
transformis Last, 1981: 132 ( Hesperus ; variety of ricardi , cited as richardi; [Note: Nomen nudum (Article 15.2)]; cited from Wantoat Valley, N. Guinea).
roepkei Bernhauer, 1911: 89 ( Hesperus ; Type locality: Java: Bandoeng; Borneo: Sarawak). — Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Gridelli, 1924: 185 ( Hesperus ; characters; Sumatra; Isole Mentawei).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 160 ( Hesperus ; characters).
- DISTRIBUTION: Borneo, Indonesia.
distincticornis Cameron, 1937: 15 ( Hesperus ; Type locality: Java: Batoerraden, G. Slamat; Bandoeng Dago; Tjilatjap; Patimoean, Zuid Preanger).
— Scheerpeltz, 1971: 160 ( Hesperus ; variety of roepkei ; characters).
rubidus Last, 1987: 44 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE: Eliptamin Valley, 1200–1350 m).
- DISTRIBUTION: New Guinea.
ruficeps Cameron, 1941c: 380 ( Hesperus ; Type locality: Philippines: Tangcolan : Bukidnon). - DISTRIBUTION: Philippines.
rufipennis Gravenhorst, 1802: 40 ( Staphylinus ; [preoccupied]; Type locality: Not cited, but presumably Brunsvigae ).
— Latreille, 1804: 327 ( Staphylinus ; characters; Germany).
— Gravenhorst, 1806: 104 ( Staphylinus ).
— Nordmann, 1837: 110 ( Philonthus ; Germany).
— Erichson, 1839a: 477 ( Philonthus ; characters; Germany).
— Erichson, 1840: 504 ( Philonthus ; characters; Germany; France; Sardinia).
— Heer, 1841: 583 ( Philonthus ; characters; Switzerland).
— Redtenbacher, 1849: 825 ( Philonthus ; characters; Austria).
— Küster, 1854: 17 ( Philonthus ; characters; Germany; France; Switzerland; Sardinia).
— Fairmaire and Laboulbène, 1856: 530 ( Philonthus ; characters; France).
— Kraatz, 1857d: 621 ( Philonthus ; characters; Germany).
— Redtenbacher, 1857: 197 ( Philonthus ; characters; Austria).
— Fauvel, 1874: 200 [= 1874a: 426] ( Hesperus ; characters; France; Germany; Austria; Italy; Sardinia).
— Redtenbacher, 1874: 215 ( Philonthus ; characters; Austria).
— Mulsant and Rey, 1876: 562 [= 1877: 418] ( Cafius ; characters; France).
— Seidlitz, 1891: 390 ( Philonthus ; subgenus Hesperus ; characters; Europe).
— Seidlitz, 1891a: 415 ( Philonthus ; subgenus Hesperus ; characters; Europe).
— Montandon, 1908: 79 ( Hesperus ; Romania).
— Reitter, 1909: 123 ( Hesperus ; characters; Germany).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Gridelli, 1924: 188 ( Hesperus ; characters; Italy; Morocco; Sardinia).
— Portevin, 1929: 355 ( Hesperus ; characters; France).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Smetana, 1958: 258 ( Hesperus ; characters; ecology; Czechoslovakia).
— Kocher, 1958: 152 ( Hesperus ; Morocco).
— Lohse, 1964: 173 ( Hesperus ; characters; central Europe).
— Smetana, 1964c: 168 ( Hesperus ; Czechoslovakia).
— Horion, 1965: 121 ( Hesperus ; France; Switzerland; Italy; Sardinia; Hercegovina; Serbia; Romania; Czech Republic; Slovakia; Poland; Germany; Austria).
— Scheerpeltz, 1968a: 79 ( Hesperus ; catalog; Austria).
— Scheerpeltz, 1971: 154 ( Hesperus ; characters).
— Coiffait, 1974: 321 ( Hesperus ; characters; Europe; Italy; Sardinia; Morocco).
— Burakowski, Mroczkowski, and Stefańska, 1980: 33 ( Hesperus ; catalog; Poland).
— Tóth, 1984: 41 ( Hesperus ; characters; Hungary).
— Lucht, 1987: 104 ( Hesperus ; checklist; Germany; Poland; Czechoslovakia; Austria; France).
— Schülke and Uhlig, 1988: 11 ( Hesperus ; notes; Germany).
— Gusarov, 1989: 11 ( Hesperus ; habitat notes; Ukraine).
— Drugmand and Coulon, 1993: 239 ( Hesperus ; collecting notes; Luxembourg).
— Smetana, 1993b: 48 ( Hesperus ; Czech Republic; Slovakia).
— Ciceroni and Zanetti, 1995: 27 ( Hesperus ; Italy).
— Herman, 2001: 45 ( Hesperus ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).
- DISTRIBUTION: Europe, Morocco.
brunneipennis Scheerpeltz, 1971: 154 ( Hesperus ; aberration of rufipennis ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from nördlichere Teile des Verbreitungsgebietes). rufipennis Cameron, 1918 , see: borneensis Bernhauer, 1915 .
sarawakensis Gridelli, 1924 , see: laevigatus Fauvel, 1895 .
saltus Last, 1968: 353 ( Philonthus ; Type locality: New Guinea: Okapa).
— Last, 1973: 425 ( Hesperus ).
— Last, 1987: 50 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
schedli Scheerpeltz, 1956b: 22 , 27 ( Hesperus ; Type locality: Belgisch-Kongo: Hembe Bitale [Note: Type locality cited in Scheerpeltz, 1956a]).
— Scheerpeltz, 1956a: 485 ( Hesperus ; type locality).
- DISTRIBUTION: Congo.
scutellaris Bernhauer, 1926b: 40 ( Hesperus ; Type locality: Luzon: Mt. Makiling).
— Scheerpeltz, 1933: 1373 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 175 ( Hesperus ; characters).
- DISTRIBUTION: Philippines.
segmentarius Cameron, 1941c: 379 ( Hesperus ; Type locality: Not cited, presumably the Philippines).
- DISTRIBUTION: Philippines.
semicoeruleus Bernhauer, 1915 h: 229 ( Hesperus ; Type locality: Java: Preanger ).
— Gridelli, 1924: 200 ( Hesperus ; characters; Java).
— Scheerpeltz, 1933: 1373 ( Hesperus ; cited as semicaeruleus; catalog).
— Scheerpeltz, 1935a: 634 ( Hesperus ; cited as semicaeruleus; Indonesia).
— Scheerpeltz, 1971: 173 ( Hesperus ; cited as semicaeruleus; characters).
- DISTRIBUTION: Indonesia.
varicornis Cameron, 1937: 15 ( Hesperus ; Type locality: Java: G. Tangkoeban Prahoe, semirufus Fauvel, 1878 g: 543 ( Hesperus ; Type locality: Australie septentrionale).
— Olliff, 1887: 511 ( Hesperus ; characters; Australia).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 176 ( Hesperus ; characters).
— Bellas, Brown, and Moore, 1974 ( Hesperus ; defensive secretions).
- DISTRIBUTION: Australia.
septuosus Herman, 2001: 45 ( Hesperus ; replacement name for obscuricollis Scheerpeltz ).
- DISTRIBUTION: Philippines.
obscuricollis Scheerpeltz, 1971: 168 ( Hesperus ; [preoccupied]; Type locality: Philippinen: Luzon : Los Baños, Subuagra).
— Herman, 2001: 45 ( Hesperus ; synonym of septuosus ).
serpentinus Horn, 1884: 219 ( Philonthus ; Type locality: Kentucky).
— Bernhauer and Schubert, 1914: 355 ( Philonthus ; catalog).
— Smetana, 1995e: 490 ( Hesperus ; characters; notes; Alabama; Florida; Kentucky; North Carolina; Pennsylvania).
— Downie and Arnett, 1996: 381 ( Philonthus ; characters; USA).
- DISTRIBUTION: USA.
setosus Cameron, 1952: 251 ( Hesperus ; Type locality: New Guinea: Snow Mountains:
Moss Forest Camp, 2800 m).
- DISTRIBUTION: New Guinea.
similis Last, 1987: 45 ( Hesperus ; Type locality: Irian: Genjam, 40 km W of Jayapura, 100–200 m).
- DISTRIBUTION: New Guinea.
singularis Last, 1973: 428 ( Philonthus ; Type locality: New Guinea: Garaina, Bula Valley).
— Last, 1987: 50 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
sociabilis Last, 1987: 45 ( Hesperus ; Type locality: Papua New Guinea: Madang).
- DISTRIBUTION: New Guinea.
solivagus Last, 1981: 133 ( Hesperus ; Type locality: Lufa, Mount Michael).
- DISTRIBUTION: New Guinea.
sparsiceps Bernhauer, 1928 g: 116 ( Hesperus ; Type locality: Insel Fernando Po: Basile , 400–600 m).
— Scheerpeltz, 1933: 1374 ( Hesperus ; catalog).
— Bernhauer, 1942c: 363 ( Hesperus ; Cameroon).
— Scheerpeltz, 1956b: 21 ( Hesperus ; characters).
— Scheerpeltz, 1971: 188 ( Hesperus ; characters).
- DISTRIBUTION: Equatorial Guinea.
sparsior Bernhauer, 1928 g: 118 ( Hesperus ; Type locality: Tanganyika: Terr.: Konga).
— Scheerpeltz, 1933: 1374 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 25 ( Hesperus ; characters; Tanganyika).
— Scheerpeltz, 1971: 186 ( Hesperus ; characters).
— Scheerpeltz, 1974c: 24 ( Hesperus ; Ethiopia; Sudan).
- DISTRIBUTION: Sudan, Ethiopia, Tanzania.
sparsus Last, 1987: 45 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE: Finisterre Range , Saidor, Kaimbaui Village).
- DISTRIBUTION: New Guinea.
spectabilis Bernhauer, 1928 g: 118 ( Hesperus ; Type locality: Natal: Malvern).
— Scheerpeltz, 1933: 1374 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 25 ( Hesperus ; characters; Natal).
— Scheerpeltz, 1971: 186 ( Hesperus ; characters).
- DISTRIBUTION: South Africa.
splendoris Last, 1987: 46 ( Hesperus ; Type locality: Papua New Guinea: New Guinea NE: Mt. Amingwiwa, 3000–3400 m).
- DISTRIBUTION: New Guinea.
stehri Moore, 1958: 313 ( Hesperus ; Type locality: Ash Cave , Hocking County, Ohio).
— Smetana, 1995e: 502 ( Hesperus ; characters; notes; Arkansas; Georgia; Kansas; Ohio; Oklahoma; Tennessee).
— Downie and Arnett, 1996: 383 ( Hesperus ; characters; USA).
- DISTRIBUTION: USA.
strigiventris Eppelsheim, 1895a: 127 ( Philonthus ; Type locality: Togoland).
— Fauvel, 1903e: 242 ( Philonthus ; Cameroon; Togo; Gabon).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Bernhauer, 1915o: 363 ( Hesperus ; Belgian Congo).
— Bernhauer, 1928g: 118 ( Hesperus ; characters in key).
— Scheerpeltz, 1933: 1374 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 26 ( Hesperus ; characters; Togo; Guinea).
— Tottenham, 1962: 144 ( Hesperus ; Uganda).
— Levasseur, 1966a: 1214 ( Hesperus ; Congo; Togo; Gabon; Zaire).
— Levasseur, 1968a: 52 ( Hesperus ; Central African Republic).
— Scheerpeltz, 1971: 183 ( Hesperus ; characters).
- DISTRIBUTION: Guinea, Togo, Cameroon, Central African Republic, Gabon, Congo, Uganda.
subgrandis Last, 1987: 46 ( Hesperus ; Type locality: Irian: Sentani, 90 m).
- DISTRIBUTION: New Guinea.
subsimilis Cameron, 1933d: 390 ( Hesperus ; Type locality: Sumatra: Gunung Singgalang , 1800 m).
— Scheerpeltz, 1971: 165 ( Hesperus ; characters).
- DISTRIBUTION: Indonesia.
sumatrensis Cameron, 1933d: 390 ( Hesperus ; Type locality: Sumatra: Fort de Kock ).
— Scheerpeltz, 1971: 164 ( Hesperus ; characters).
- DISTRIBUTION: Indonesia.
taiwanensis Shibata, 1973: 21 ( Hesperus ; Type locality: Taiwan: Koantauchi, Nantou Hsien). — Shibata, 1990: 212 ( Hesperus ; Taiwan).
- DISTRIBUTION: Taiwan.
tapiniensis Last, 1987: 46 ( Hesperus ; Type locality: Papua New Guinea: New Guinea SE: Tapini, 1000 m).
- DISTRIBUTION: New Guinea.
tectus Last, 1968: 364 ( Philonthus ; Type locality: New Guinea: Buana , Sarawaged Range, 3000 ft., 20–30 miles north of Lae).
— Last, 1973: 425 ( Hesperus ; characters).
- DISTRIBUTION: New Guinea.
temporatus Last, 1968: 348 ( Philonthus ; Type locality: New Guinea: Okapa).
— Last, 1973: 425, 426 ( Hesperus ; characters).
— Last, 1987: 50 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
tenuis Last, 1973: 430 ( Hesperus ; Type locality: Okapa).
— Last, 1987: 50 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
terminalis Cameron, 1918a: 104 ( Hesperus ; Type locality: Java: Silabintanah ).
— Gridelli, 1924: 200 ( Hesperus ; characters; Java).
— Scheerpeltz, 1933: 1374 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 171 ( Hesperus ; characters).
terminicornis Gridelli, 1924: 182 , 192 ( Hesperus ; Type locality: Beccari a Sumatra,
Monte Singalang).
— Scheerpeltz, 1933: 1374 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 169 ( Hesperus ; characters).
- DISTRIBUTION: Indonesia.
tiro Sharp, 1874: 49 ( Philonthus ; Type locality: Copper Temple , Nagasaki).
— Bernhauer and Schubert, 1914: 358 ( Philonthus ; catalog).
— Scheerpeltz, 1933: 1374 ( Hesperus ; catalog).
— Nakane, 1963a: 92 ( Hesperus ; characters; habitus photograph; Japan).
— Scheerpeltz, 1971: 156 ( Hesperus ; characters).
— J. Li, 1993: 40 ( Hesperus ; China).
— Watanabe, 1996a: 12 ( Hesperus ; mine collection; Japan).
- DISTRIBUTION: Japan, China.
tokyanus R. Dvořák, 1957: 9 ( Hesperus ; Type locality: Mount Takar , Tokyo, Japon).
- DISTRIBUTION: Japan.
toxopei Cameron, 1952: 253 ( Hesperus ; Type locality: New Guinea: Snow Mountains: Rattan Camp, 1200 m).
- DISTRIBUTION: New Guinea.
transformis Last, 1981 [Note: Nomen nudum], see: ricardi Last, 1973 .
tremolerasi Bernhauer, 1927e: 246 ( Hesperus ; Type locality: Uruguay: Montevideo).
— Scheerpeltz, 1933: 1374 ( Hesperus ; catalog).
— Scheerpeltz, 1936a: 490 ( Hesperus ; characters in key).
— Scheerpeltz, 1971: 191 ( Hesperus ; characters).
- DISTRIBUTION: Uruguay.
tricolor Cameron, 1952: 251 ( Hesperus ; Type locality: New Guinea: Snow Mountains : Moss Forest Camp, 2800 m).
— Last, 1987: 50 ( Hesperus ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
trimerus Fauvel, 1895: 261 ( Hesperus ; Type locality: Bírmanie, Carin Cheba, 900–1100 m). — Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Gridelli, 1924: 189 ( Hesperus ; characters; Burma).
— Cameron, 1932: 159 ( Hesperus ; characters; Burma; India).
— Scheerpeltz, 1933: 1374 ( Hesperus ; catalog).
— Scheerpeltz, 1965: 108 ( Hesperus ; Burma).
— Scheerpeltz, 1971: 158 ( Hesperus ; characters).
- DISTRIBUTION: Myanmar, India.
uelensis Bernhauer, 1932b: 153 ( Hesperus ; Type locality: Haut-Uele: Moto).
— Scheerpeltz, 1933: 1374 ( Hesperus ; catalog).
— Scheerpeltz, 1956b: 25 ( Hesperus ; characters; Belgian Congo).
— Scheerpeltz, 1971: 181 ( Hesperus ; characters).
- DISTRIBUTION: Congo.
uhligi Last, 1990: 1 ( Hesperus ; Type locality: Deutsch Neu Guinea: Mäanderberg, 470 m). - DISTRIBUTION: New Guinea.
varicornis Cameron, 1937 , see: semicaeruleus Bernhauer, 1915.
venator Last, 1987: 48 ( Hesperus ; Type locality: Irian: Vogelkop, Bomberai, 700–900 m).
- DISTRIBUTION: New Guinea.
venustulus Last, 1968: 351 ( Philonthus ; Type locality: New Guinea: Okapa).
— Last, 1973: 425, 426 ( Hesperus ; characters).
- DISTRIBUTION: New Guinea.
venustus Last, 1968: 350 ( Philonthus ; Type locality: New Guinea: Agakomatuso).
— Last, 1973: 425 ( Hesperus ).
vethi Bernhauer, 1915 h: 230 ( Hesperus ; Type locality: Java: Preanger ).
— Gridelli, 1924: 191 ( Hesperus ; characters; Java).
— Scheerpeltz, 1933: 1374 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 165 ( Hesperus ; characters).
- DISTRIBUTION: Indonesia.
viridiaureus Bernhauer, 1926b: 41 ( Hesperus ; variety of phaenomenalis ; Type locality: Mindanao: Surigao; Luzon: Imugan).
— Scheerpeltz, 1933: 1374 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 175 ( Hesperus ; characters).
- DISTRIBUTION: Philippines.
viridipennis Cameron, 1952: 249 ( Hesperus ; Type locality: Misool Island).
- DISTRIBUTION: Indonesia.
viridis Bernhauer, 1926 , see: phaenomenalis Bernhauer, 1914 .
viridisplendens Scheerpeltz, 1971: 175 ( Hesperus ; Type locality: Philippinen: S. O. Luzon: Mt. Balusan ).
- DISTRIBUTION: Philippines.
vividus Cameron, 1931 , see: Philonthus .
wasmanni Fauvel, 1895: 260 ( Hesperus ; Type locality: Bírmanie, Carin Cheba , 900–1100 m; Sikkim, Darjeeling).
— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).
— Gridelli, 1924: 187 ( Hesperus ; characters; Burma; India).
— Cameron, 1932: 158 ( Hesperus ; characters; Burma; India).
— Scheerpeltz, 1933: 1374 ( Hesperus ; catalog).
— Scheerpeltz, 1971: 155, 158 ( Hesperus ; characters).
- DISTRIBUTION: Myanmar, India.
zischkai Scheerpeltz, 1971: 192 ( Hesperus ; Type locality: Süd-ost-Bolivien in der Sierra de Cochabamba).
- DISTRIBUTION: Bolivia.
[8 species; Palaearctic Region]
Hybridolinus Schillhammer, 1998 a: 146 (species included: hesperoides, singularis, diabolicus, daliensis, similis, jizushanus, decipiens). TYPE SPECIES: Hybridolinus daliensis Schillhammer, fixed by original designation.
— Schillhammer, 2000 b: 144 (key to species).
— Hayashi, 2000: 23 (notes on empodial setae).
daliensis Schillhammer, 1998 a: 149 (Hybridolinus; Type locality: China: Yunnan, Dali).
— Schillhammer, 2000 b: 144 (Hybridolinus; characters in key).
- DISTRIBUTION: China.
decipiens Schillhammer, 1998 a: 150 (Hybridolinus; Type locality: China: Yunnan, Baishui).
— Schillhammer, 2000 b: 144 (Hybridolinus; characters in key).
- DISTRIBUTION: China.
diabolicus Schillhammer, 1998 a: 154 (Hybridolinus; Type locality: Taiwan: Taichung Hsien, Anmashan, 2230 m).
— Schillhammer, 2000 b: 144 (Hybridolinus; characters in key).
- DISTRIBUTION: Taiwan.
flavocinctus Schillhammer, 2000 b: 143 (Hybridolinus; Type locality: China: SE Sichuan: Jinfo Shan, 29° 01N, 107°14E, 1750 m).
- DISTRIBUTION: China.
hesperoides Schillhammer, 1998 a: 154 (Hybridolinus; Type locality: China: Kuatun, 2300 m).
— Schillhammer, 2000 b: 144 (Hybridolinus; characters in key).
jizushanus Schillhammer, 1998 a: 152 (Hybridolinus; Type locality: China: Yunnan, Jizu-Shan).
— Schillhammer, 2000 b: 144 (Hybridolinus; characters in key).
- DISTRIBUTION: China.
similis Schillhammer, 1998 a: 150 (Hybridolinus; Type locality: China: Sichuan, Daxue Shan, Gongga Shan Mt., Hailuogou glacier park, 102.04 E, 29.36 N, river valley ca. 1 km above Camp 1, 2100 m).
— Schillhammer, 2000 b: 144 (Hybridolinus; characters in key).
- DISTRIBUTION: China.
singularis Schillhammer, 1998 a: 152 (Hybridolinus; Type locality: China: Yunnan, Jizu-Shan). — Schillhammer, 2000b: 144 (Hybridolinus; characters in key).
- DISTRIBUTION: China.
[2 species; Palaearctic Region]
Jurecekia Rambousek, 1920: 16 , 19 (species included: paradoxa ). TYPE SPECIES: Jurecekia paradoxa Rambousek , fixed by monotypy.
— Winkler, 1925: 381 (catalog for Palaearctic region).
— Cameron, 1932: 62 (subgenus of Philonthus ).
— Scheerpeltz, 1933: 1371 (world catalog supplement).
— Blackwelder, 1952: 206 (type species: paradoxa ).
— Tikhomirova, 1973a: 182 (checklist of species of USSR).
— Coiffait, 1974: 316 (characters; key to species of western Palaearctic region).
— Smetana, 1995e: 50 (characters).
asphaltina Erichson, 1840: 436 ( Philonthus ; Type locality: Germania meridionalis [Note: Erroneous type locality, see Horion, 1965: 138]).
— Kraatz, 1857d: 576 ( Philonthus ; characters; Germany).
— Redtenbacher, 1874: 216 ( Philonthus ; characters).
— Ganglbauer, 1895: 445 ( Philonthus ; characters; southern Germany).
— Bernhauer, 1899a: 434 ( Philonthus ; characters; Austria).
— Reitter, 1909: 126 ( Philonthus ; characters; Germany).
— Bernhauer and Schubert, 1914: 329 ( Philonthus ; catalog).
— Scheerpeltz, 1933: 1371 ( Jurecekia ; catalog).
— Horion, 1965: 138 ( Jurecekia ; species not found in central Europe).
— Coiffait, 1974: 317 ( Jurecekia ; characters; Russia; Transcaucasus; Germany).
- DISTRIBUTION: Russia, Transcaucasus.
paradoxa Rambousek, 1920: 17 , 19 ( Jurecekia ; Type locality: Russie aux environs de Totskoe, (Gouber. Samara) au bord de Samarka).
— Rambousek, 1923: 54 ( Philonthus ; synonym of asphaltinus ).
— Scheerpeltz, 1933: 1371 ( Jurecekia ; synonym of asphaltina ).
— Coiffait, 1974: 318 ( Jurecekia ; synonym of asphaltina ).
paradoxa Rambousek, 1921 , see: asphaltina Erichson, 1840 .
rufipes Coiffait, 1969b: 39 ( Jurecekia ; Type locality: Atlosar région d’Akmolinsk, Kazakstan). — Coiffait, 1974: 319 ( Jurecekia ; characters; Kazakhstan).
- DISTRIBUTION: Kazakhstan.
[2 species; Nearctic Region]
Laetulonthus Moore and Legner, 1973c: 213 (species included: laetulus ). TYPE SPECIES: Staphylinus laetulus Say , fixed by original designation.
— Moore and Legner, 1975: 17 (catalog; North America north of Mexico).
— Moore and Legner, 1979: 23 (characters; notes).
— Smetana, 1995e: 505 (characters; notes; phylogenetic relationships; key to species). — Downie and Arnett, 1996: 383 (characters). — Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species). — Newton, Thayer, Ashe, and Chandler, 2000: 396 (2 Nearctic species; notes; characters in key).
laetulus Say, 1834: 449 ( Staphylinus ; [Note: This species was described in a note within a discussion of blandus Gravenhorst ]; Type locality: Not cited [Note: See neotype designation by Smetana, 1995e]). — Melsheimer, 1844: 36 ( Philonthus ; characters; Pennsylvania). — LeConte, 1859: 564 ( Philonthus ). — Horn, 1884: 184 ( Philonthus ; characters; Canada to Georgia west to the Mississippi). — Blatchley, 1910: 381 ( Philonthus ; characters; Indiana). — Bernhauer and Schubert, 1914: 343 ( Philonthus ; catalog). — Moore and Legner, 1973c: 215 ( Philonthus ; characters; notes). — Legner and Moore, 1977: 176 ( Laetulonthus ; found under bark). — Frank, 1984: 478 (Laetulontus; notes; Florida). — Campbell and Davies, 1991: 119 ( Laetulonthus ; checklist of species; Canada). — Smetana, 1995e: 508 ( Laetulonthus ; neotype designation: [Note: Locality not cited; specimen selected from LeConte collection with “yellow round disc”]; characters; notes; Ontario; Quebec; Alabama; Arkansas; Connecticut; Florida; Georgia; Illinois; Indiana; Kentucky; Louisiana; Maine; Maryland; Massachusetts; Michigan; Mississippi; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; South Carolina; Texas; Vermont; Virginia; West Virginia; Wisconsin). — Downie and Arnett, 1996: 383 ( Laetulonthus ; characters; USA). - DISTRIBUTION: Canada, USA.
nobilis Bernhauer, 1910b: 381 ( Philonthus ; Type locality: Neu Mexico: Porvenir). — Bernhauer and Schubert, 1914: 348 ( Philonthus ; catalog). — Smetana, 1995e: 512 ( Laetulonthus ; characters; notes; Arizona; New Mexico). - DISTRIBUTION: USA.
[3 species; Neotropical Region]
Leptopeltus Bernhauer, 1906c: 337 (species included: flavipennis , netolitzkyi , montivagans ). TYPE SPECIES: Philonthus flavipennis Erichson , fixed by subsequent designation by R. Lucas, 1920: 371.
— Bernhauer and Schubert, 1914: 365 (world catalog; 3 species).
— Bruch, 1928a: 428 (catalog; Argentina).
— Scheerpeltz, 1933: 1374 (world catalog supplement).
— Blackwelder, 1944: 138 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 217 (type species).
— Scheerpeltz, 1960c: 114 (key to species).
flavipennis Erichson, 1840: 460 ( Philonthus ; Type locality: Bogotae in Columbia).
— Solsky, 1872: 299 ( Philonthus ; Colombia).
— Bernhauer, 1906c: 338 ( Leptopeltus ).
— Bernhauer and Schubert, 1914: 365 ( Leptopeltus ; catalog).
— Scheerpeltz, 1933: 1374 ( Leptopeltus ; catalog; Bolivia, Peru, Ecuador, Venezuela).
— Scheerpeltz, 1960c: 115 ( Leptopeltus ; characters; Venezuela to Chile).
- DISTRIBUTION: Venezuela, Colombia, Ecuador, Peru, Bolivia.
montivagans Bernhauer, 1906c: 339 ( Leptopeltus ; Type locality: Bolivia: Hochanden ,
4000 m).
— Bernhauer and Schubert, 1914: 365 ( Leptopeltus ; catalog).
— Scheerpeltz, 1960c: 115 ( Leptopeltus ; characters; Bolivia).
netolitzkyi Bernhauer, 1906c: 338 ( Leptopeltus ; Type locality: Bolivia: La Paz). — Bernhauer and Schubert, 1914: 365 ( Leptopeltus ; catalog). — Scheerpeltz, 1960c: 115 ( Leptopeltus ; characters; Bolivia). - DISTRIBUTION: Bolivia.
[12 species; Oriental, Australian, and Oceanic Regions]
Leucitus Fauvel, 1878f: 253 (species included: stenoides [with doubt] and argyreus ). TYPE SPECIES: Leucitus argyreus Fauvel , fixed by virtual monotypy.
— Duvivier, 1883: 152 (catalog).
— Bernhauer and Schubert, 1914: 372 (world catalog; 8 species).
— Scheerpeltz, 1933: 1380 (world catalog supplement).
— Blackwelder, 1952: 219 (type species: argyreus ).
— Last, 1975a: 1 (characters; type species designation).
— Hammond, 1984: 206 (checklist of species of Borneo).
amicus Bernhauer, 1915 g: 193 ( Leucitus ; Type locality: Neu-Guinea: Sekar).
— Scheerpeltz, 1933: 1380 ( Leucitus ; catalog).
— Last, 1975a: 3 ( Leucitus ; characters).
- DISTRIBUTION: New Guinea.
argyreus Fauvel, 1878f: 254 ( Leucitus ; Type locality: Amberbaki , New Guinea; [Note: See lectotype designation by Last, 1975a]. Also cited from Nouvelle-Guinée, Ramoi; Iles Arou, Wokan; Mysol).
— Fauvel, 1879: 96 ( Leucitus ; characters; notes; Dorei; Vaigheu).
— Heller, 1910: 10 ( Leucitus ; characters in key).
— Bernhauer and Schubert, 1914: 372 ( Leucitus ; catalog).
— Arrow, 1915: 500 ( Leucitus ; Dutch New Guinea).
— Cameron, 1924: 87 ( Leucitus ; New Guinea).
— Cameron, 1931: 363 ( Leucitus ; New Guinea).
— Last, 1975a: 3 ( Leucitus ; lectotype designation: Amberbaki , New Guinea; characters) .
— Last, 1987: 55 ( Leucitus ; Papua New Guinea).
- DISTRIBUTION: New Guinea, Indonesia.
benigseni Bernhauer, 1927 , see: Philonthus .
bernhaueri Heller, 1910: 10 ( Leucitus ; Type locality: Nova Guinea britannica, Astrolabe montes).
— Bernhauer and Schubert, 1914: 373 ( Leucitus ; catalog).
— Last, 1975a: 4 ( Leucitus ; characters).
— Last, 1987: 55 ( Leucitus ; Papua New Guinea; Irian).
- DISTRIBUTION: New Britain, New Guinea.
biroi Last, 1980: 155 ( Leucitus ; Type locality: New Guinea: Sattelberg, Huon Golf).
- DISTRIBUTION: New Guinea.
freudei Last, 1975a: 5 ( Leucitus ; Type locality: Papua / Wau, Keinde ).
- DISTRIBUTION: New Guinea.
mandibularis Heller, 1910: 9 ( Leucitus ; Type locality: Papua New Guinea, 780 m).
— Bernhauer and Schubert, 1914: 373 ( Leucitus ; catalog).
— Last, 1975a: 4 ( Leucitus ; characters).
- DISTRIBUTION: New Guinea.
paradiseus Fauvel, 1879: 96 ( Leucitus ; Type locality: Nouva Guinea, Fly River,
Mt. Hautukeen ; [Note: See lectotype designation by Last, 1975a]. Originally cited from Nouvelle Guinée, Fly River ).
— Bernhauer and Schubert, 1914: 373 ( Leucitus ; catalog).
— Last, 1975a: 4 ( Leucitus ; lectotype designation: Nouva Guinea, Fly River, Mt. Hautukeen; characters) .
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