Pipunculus violovitshi Kuznetzov, 1991, Kuznetzov, 1991

Kehlmaier, Christian, 2008, Finnish Pipunculidae (Diptera) studies Part I: Taxonomic notes on Cephalops Fallén, 1810, Pipunculus Latreille, 1802 and Tomosvaryella Aczél, 1939, Zootaxa 1672, pp. 1-42: 26-27

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http://doi.org/ 10.5281/zenodo.180216

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Pipunculus violovitshi Kuznetzov, 1991


Pipunculus violovitshi Kuznetzov, 1991  

( Figs 1, 5 View FIGURES 1 – 8 , 43 –44 View FIGURES 43 – 54 , 74 View FIGURES 74 – 83 & 84–85 View FIGURES 84 – 92 )

Pipunculus varipes   auct. nec Meigen, 1824: 21 Pipunculus violovitshi Kuznetzov, 1991: 2  

Material: Finland, N, Tvärminne village, 6642: 287: 1 ɗ, 14.– 27.VI. 1990; 1 ɗ, 1.– 15.VII. 1990, Albrecht; Finland, N, Sibbo, Nevas, 6687: 412: 1 ɗ, 10.– 15.VII. 1989; 1 ɗ, 5.– 11.VII. 1987, Albrecht; Germany, Bavaria, Oberallgäu, S of Fischen, 10 ° 16 ’ 42 ’’E 47 ° 27 ’ 11 ’N, 760m: 1 Ψ, 15.VIII. 2004, Kehlmaier ( PCCK); Germany, Sachsen, Weißeritzkreis, Kurort Kipsdorf, NSG Hofehübel, 13 ° 40 ’06’’E 50 ° 48 ’09’’N, 670m: 1 ɗ 1 Ψ, 5.VII. 2002, Kehlmaier ( PCCK).

Male: Body length: 5.0– 5.6mm. Wing length: 5.4–6.1mm. Flagellum smaller than in other species and dark brown ( Fig. 84 View FIGURES 84 – 92 ). Frons grey pollinose. Occiput entirely grey pollinose but brown pollinosity can be present posterior to ocellar triangle ( Fig. 84 View FIGURES 84 – 92 ). Eyes meeting for slightly less than length of frons (14–16 facets). Postpronotal lobe dark brown. Prescutum and scutum weakly grey to brown pollinose in anterior fifth to quarter, narrowly along lateral and posterior margin (occasionally also weakly dorsocentrally), otherwise shining. Pleura with anepimeron weakly grey to brown pollinose, otherwise with weak grey pollinosity. Scutellum with apical fringe of about 20 long hairs, the longest ones about half of its maximum length ( Fig. 74 View FIGURES 74 – 83 ). Subscutellum grey pollinose. LTC about 1.2–1.9 times LFC. Trochanter yellow. Femur dark brown with base narrowly to basal quarter and apical quarter to third yellow. Hind femur evenly and gently covex ventrally. Tibia yellow, can be slightly darkened ventromedially. Tarsi yellow, with distitarsi darkened. Pulvilli of front legs slightly shorter than distitarsi. Tergite 1 anterodorsally brown, otherwise grey pollinose. Tergites 2– 5 laterally shining or narrowly and weakly greyish-brown pollinose, dorsally with brown pollinosity in anterior two thirds (tergite 2), half (tergite 3) or only narrowly (tergites 4–5). Membranous area narrow, slot-like, broadest in lower part ( Fig. 85 View FIGURES 84 – 92 ). Surstyli narrow in dorsal and almost straight in lateral view ( Figs 1 & 5 View FIGURES 1 – 8 ). Phallus trifid with ejaculatory ducts almost circular and conspicuously sclerotized, dark brown (in one occasion almost hyaline in distal half).

Female: Body length: 4.3–5.3mm. Wing length: 5.3–5.9mm. Flagellum mid to dark brown and smaller than in other species. Frons entirely grey pollinose. Postpronotal lobe yellow. Prescutum and scutum grey pollinose in anterior quarter, broadly along lateral and narrowly along posterior margin, centrally shining. LTC about 0.9 –1.0 times LFC. Pleura and subscutellum grey pollinose. Trochanter yellow. Femur with basal quarter or less and apical third yellow. Tibia yellow, darkened ventromedially. Tarsi yellow, distitarsi darkened. Pulvilli of front leg as long as distitarsi. Tergite 1 brown pollinose in anterodorsal half, otherwise with grey pollinosity. Tergites 2–6 laterally grey pollinose. Dorsally, tergite 2 grey pollinose in anterior half, otherwise shining like tergites 3–6. Ovipositor in lateral view with strongly curved piercer, showing a distinct ventral step ( Fig. 43 View FIGURES 43 – 54 ).

Annotations: Due to the misinterpretation of P. v a r i p e s Meigen, 1824 by all former authors and the subsequent transfer to Cephalops   (see above), the specimens formerly addressed as Pipunculus varipes   auct. nec Meigen, 1824 (in the following abbreviated as “ varipes   ”) are left without any available name. It is proposed to address them as Pipunculus violovitshi Kuznetzov, 1991   for the time being due to the following reasons:

• The description of Pipunculus violovitshi Kuznetzov, 1991   is based on two males from Sakhalin Island, Russian Far East, deposited at ZISP and, although attempted, could not be obtained for study. According to Kuznetzov (1991: 2), P. v i o l o v i t s h i differs from “ varipes   ” “… by the small third antennal segment and considerably prolonged surstyli of the male genitalia ”. Judging from his figures, the surstyli only seem to be very slightly longer compared to the specimens studied here and the flagellum, not figured by Kuznetzov, is found to be generally somewhat smaller in “ varipes   ” compared to other European taxa of Pipunculus   . Therefore, Kuznetzov’s P. violovitshi   is regarded conspecific with P. v a r i p e s auct. nec Meigen, 1824.

Pipunculus monticola Schummel, 1837   , synonymized with P. v a r i p e s by Becker (1921: 156), is regarded as a nomen dubium here (see above) and hence, cannot be applied to “ varipes   ”. Verrall (1901: 102 f) quotes the brief original description of this taxon. The whereabouts of the single type specimen is unknown at present and no subsequent writer seems to have been able to examine it.

• Morakote & Hirashima (1990) investigated the Japanese Pipunculus   fauna and introduced Pipunculus subvaripes Morakote, 1990   and Pipunculus magnicarinatus Morakote, 1990   two species—both with a male holotype—closely allied to “ varipes   ”, which they also record from Japan. Note that the female ovipositor of P. subvaripes   figured (their Figs. 57 View FIGURES 55 – 66 A, C) is more like “ varipes   ” whereas P. v a r i p e s sensu Morakote & Hirashima (1990) (their Fig. 57 View FIGURES 55 – 66 B) is distinctly different, which might indicate an incorrect association of the sexes. However, male P. v a r i p e s sensu Morakote & Hirashima (1990) (their Figs 54 View FIGURES 43 – 54 A, B) corresponds with the concept of P. v a r i p e s auct. nec Meigen, 1824 whereas P. subvaripes   and P. magnicarinatus   differ in coloration and genitalic features, following the original description.


Zoological Institute, Russian Academy of Sciences














Pipunculus violovitshi Kuznetzov, 1991

Kehlmaier, Christian 2008

Pipunculus varipes

Kuznetzov 1991: 2
Meigen 1824: 21