Selenocopris Burmeister

Nunes, Rafael V. & Vaz-De-Mello, Fernando Z., 2016, New brachypterous species of Dichotomius (Selenocopris) Burmeister (Coleoptera: Scarabaeidae: Scarabaeinae) with the definition of species groups and taxonomic notes in the subgenus, Zootaxa 4139 (1) : -

publication ID

https://doi.org/ 10.11646/zootaxa.4139.1.4

publication LSID

lsid:zoobank.org:pub:2744EC46-70A1-4267-AAB8-2131F5DC21DC

DOI

https://doi.org/10.5281/zenodo.6090767

persistent identifier

https://treatment.plazi.org/id/03C187AE-453A-FF83-0EC8-F8AEFA00F94C

treatment provided by

Plazi

scientific name

Selenocopris Burmeister
status

 

Key for species groups and brachypterous species of Selenocopris Burmeister View in CoL

The following key allows the identification of the Selenocopris species groups and the brachypterous species of the subgenus, now assigned to the quadraticeps species group. When not specified, the characters fit both for males and females. Females differ from males by always having the 6th abdominal sternite not shortened medially (usually three times wider than the 5th sternite) and/or having modifications on the median portion of it or at the apex of pygidium ( Figs. 1 View FIGURE 1 A, 1H–L).

1 Clypeo-genal junction border rounded, lacking angulation ( Fig. 1 View FIGURE 1 F) and/or clypeal teeth separated from adjacent clypeal bor- der by external emarginations ( Fig. 1 View FIGURE 1 F). Female pygidium swollen, apex excavated in some species ( Figs. 2 View FIGURE 2 B–C) and/or female 6th abdominal sternite bearing a rounded central lobe advancing above pygidium apex. ( Fig. 1 View FIGURE 1 I). South portion of Atlantic Forest ( Brazil, Paraguay, Argentina) and gallery forests at Central Brazil and South America Chaco ................................................................................................. bicuspis View in CoL species group

- Clypeo-genal junction border not rounded, forming at least an obtuse angle, which may be very obtuse on smaller males and females ( Fig. 1 View FIGURE 1 G), clypeal teeth never separated from adjacent clypeal border by external emarginations ( Fig. 1 View FIGURE 1 G). Female pygidium normally convex, apex never excavated; female 6th abdominal sternite either 3x wider than other segments ( Fig. 1 View FIGURE 1 L) or having other modifications at the median portion (emarginations, grooves, projecions or tubercles) ( Figs. 1 View FIGURE 1 H, 1K). Forest habitats in South America............................................................................... 2

2 Male 6th abdominal ventrite with strong transverse groove all along its extension ( Fig. 1 View FIGURE 1 J). Female 6th abdominal ventrite with a "U" shape emargination at medial portion ( Fig. 1 View FIGURE 1 K). Apex of male paramera curved inward (dorsal and ventral view) lacking rounded or circular excavations near basis (dorsal view). Pronotum simply convex, having at most a pair of tubercles at anterior portion......................................................................... ascanius View in CoL species group

- Male 6th abdominal segment lacking strong transverse groove all over its extension. Female 6th abdominal sternite with tubercles or other modifications at medial portion ( Figs. 1 View FIGURE 1 A,H,L). Male paramera curved inward or outward (dorsal and ventral view) and/or truncated at apex and having rounded or circular excavations near basis (dorsal view) ( Fig. 3 View FIGURE 3 C–D). Pronotum simply convex or having excavations, tubercles, lobes or strong declivity at anterior portion......................... 3

3 Male clypeo-genal junction border either in straight sub-straight or even acute angle, in some cases expanded ( Fig. 1 View FIGURE 1 G). Genal border usually curved inward and clypeal teeth always curved upward. Female 6th abdominal sternite or 3x larger then 5th segment medially or having a central pair of tubercles ( Figs. 1 View FIGURE 1 A, 1L)...... quadraticeps View in CoL species group (continues on couplet 5)

- Male clypeo-genal junction border always obtuse and never expanded. Genal border straight and clypeal teeth curved upward or not. Female 6th abdominal sternite with a pair of triangular projections advancing beneath pygidial apex ( Fig. 1 View FIGURE 1 H)..... 4

4 Body almost always over 15 mm in length. Elytral striae and its punctures shallowly and weakly impressed ( Fig. 1 View FIGURE 1 D). Elytral interstriae flat, with either silky aspect given by its microsculpture (chagrination) and/or bearing brown or blue reflections. Male paramera bearing longitudinal excavations (dorsal and ventral view) (as those on Fig. 3 View FIGURE 3 C–D)...... fissus View in CoL species group

- Body almost always under 15 mm in length. Elytral striae and its punctures deeply impressed ( Fig. 1 View FIGURE 1 E). Elytral interstriae shinny, not bearing microsculpture (chagrination), sometimes with color reflections. Male paramera lacking longitudinal excavations.......................................................................... spadiceus View in CoL species group

5 Brachypterous species (atrophied hind wings) ( Fig. 2 View FIGURE 2 E): lacking humeral callus; pronotum wider than elytra (dorsal view); elytra strongly convex (lateral view) ( Fig. 1 View FIGURE 1 B).............................................................. 6

- Macropterous species (fully developed hind wings) ( Fig. 2 View FIGURE 2 E): having humeral callus; pronotum with the same width as elytra (dorsal view); elytra normally convex (lateral view).............................. macropterous species not dealt here

6 Pygidium bearing ocellate punctures near anterior margin (basis) ( Fig. 2 View FIGURE 2 D). Chapada dos Guimarães, Mato Grosso state, Brazil........................................................... Dichotomius (S.) ingens (Luederwaldt 1935) View in CoL ( Fig. 3 View FIGURE 3 )

- Pygidium lacking ocellate punctures ( Fig. 2 View FIGURE 2 C), at most having very fine punctures (20x)............................. 7

7 Posterior portion of pronotal disc bearing ocellate punctures ( Figs. 6 View FIGURE 6 A–B). Elytral striae widened, lacking distinct punctures ( Figs. 6 View FIGURE 6 A–B, E). Chapada dos Parecis at Mato Grosso state, Brazil................ Dichotomius (S.) paresi n.sp. ( Fig. 6 View FIGURE 6 )

- Pronotal disc lacking ocellate punctures, those being sometimes restricted to posterior margin, anterior angles and/or to the anterior excavation. Elytral striae narrow and always bearing distinct punctures all over its extension ( Fig. 1 View FIGURE 1 E).......... 8

8 Cephalic carina surface wrinkled (striated) ( Figs. 4 View FIGURE 4 A–B, E). Anterior excavation of pronotum bearing coarse punctures ( Fig. 4 View FIGURE 4 F). Chapadão do Céu, Goiás state, Brazil.............................................. Dichotomius (S.) darwini View in CoL

- Cephalic carina lacking striation. Anterior excavation of pronotum bearing at most fine punctures (20 x) (as on Fig. 7 View FIGURE 7 E)... 9

9 Clypeo-frontal horn carinate, strongly transverse, widely emarginated at the apex and producing two tubercles at each apex ( Fig. 7 View FIGURE 7 A). On smaller individuals, cephalic horn not producing lateral tubercles but still with a conspicuous emargination at apex. Smaller species (maximum body length: 19.5 mm; maximum pronotum width: 13 mm). Descalvado Municipality, São Paulo state, Brazil..................................................... Dichotomius (S.) periotoi View in CoL n. sp. ( Fig. 7 View FIGURE 7 )

- Clypeo-frontal horn carinate but not strongly transverse, lacking emargination and tubercles at the apex ( Fig. 5 View FIGURE 5 A). Females and smaller individuals unknown. Larger species (maximum body length: 25 mm; maximum pronotum width: 14 mm). Serra da Canastra, Southwest of Minas Gerais state, Brazil.............................. Dichotomius (S.) manni View in CoL n. sp. ( Fig. 5 View FIGURE 5 )

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

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