Winnertzia ombergensis, Jaschhof & Jaschhof, 2020

Jaschhof, Mathias & Jaschhof, Catrin, 2020, Reevaluation of species richness in Winnertzia (Diptera, Cecidomyiidae, Winnertziinae), with descriptions of 37 new species from Sweden, Peru and Australia, Zootaxa 4829 (1), pp. 1-72 : 50-51

Winnertzia ombergensis sp. nov.

Figs 85–89

Diagnosis. As discussed under W. bulbifera , W. ombergensis belongs to a group of species with long, inflated gonostyli and markedly bulging medial gonocoxal bridges (the bulbifera subgroup of the solidaginis group). This large, brown Winnertzia is distinguished from generally similar species as follows. The swollen portion of the gonostylus is situated subapically, followed by a short tapered portion ( Fig. 86 , ↓ 4); the broad pectinate claw occupies the full width of the gonostylar apex (↓ 5). The ninth tergite is unique in that its posterior portion consists of two slightly protruding, setose lobes laterally, which are interlinked by a narrow, unsclerotized portion medially ( Fig. 85 , ↓ 6). The gonocoxal synsclerite, which is markedly longer than that found in W. bulbifera agg., is strongly narrowed towards the base. Females and preimaginal stages of W. ombergensis are unknown.

Other male characters. Body size 2.7 mm. Head. Eye bridge 4–5 ommatidia long dorsally. Antenna two thirds body length. Scape slightly larger than pedicel, both concolorous with flagellum. 12 flagellomeres, all with translucent sensilla. Fourth flagellomere: neck 0.8 times as long as node; node 1.8 times as long as broad; sensory hairs numerous; both lateral and medial translucent sensilla obliquely aligned, filiform to slightly broadened, mostly variously U-shaped ( Figs 88–89 ). Palpus as long as head height, 4 setae-bearing segments; fourth segment slightly longer than third. Labella of normal size. Thorax. Pronotal setae 20–25. Anepimeral setae absent. Lateral mediotergal microtrichia strongly enlarged. Parascutellar area bright, sharply contoured. Wing as long as body, 2.6 times as long as broad. Costal cell reinforced. M 4 long, gently bent just as CuA, both veins extending to edge of wing. Legs. Scales pointed. Basitarsal spines absent. Fore tibia 0.9 times length T 2. Acropods: claws slightly bent, basal tooth large; empodia claw-long. Abdomen. Pleural membrane setose. Genitalia ( Fig. 87 ). Ninth tergite two thirds gonopodal length; anterior edge straight, distinct. Gonocoxal synsclerite broader than long; a large portion ventrobasally asetose; ventral emargination shaped like a widely open U, accompanied basally by extensive unsclerotized area; ventroanterior edge indistinct, convex; ventro- and dorsoposterior portions ending at same level; dorsal apodemes long. Gonostylus 2.5 times as long as broad, markedly convex posteriorly, very slightly convex medially; basolateral apophysis small, angulated. Aedeagal apodeme gently broadened towards apex, then suddenly constricted; solid basal portion short. Aedeagal bulge with closely spaced rows of tiny spikes. Tegmen elongatesubtriangular, largely membranous, vaguely contoured except for basal half; flaps large, faintly contoured, without microtrichia; parameral apodemes large.

Etymology. The Omberg, from which the specific name is derived, is a horst mountain in Östergötland, where the type specimens were collected.

Type material. Holotype. Male, Sweden, Östergötland, Ödeshög, Omberg , Storpissan Nature Reserve , oldgrowth forest of Norway spruce, 6 June 2010, aspirator, M. & C. Jaschhof (spn GULI000021265 in NHRS) . Paratype. 1 male, same data as the holotype (spn GULI000021265 in NHRS) .

Remark. Both specimens were referred to in our 2013 treatise on Winnertzia under W. nigripennis ( Jaschhof & Jaschhof 2013: 92) , a misidentification that is herewith corrected.

Distribution and phenology. The types, whose collection data are specified above, are the only specimens known of this species.

Figures

FIGURES 84–89. Male morphology of Winnertzia lobata (84) and Winnertzia ombergensis (85–89), holotypes. 84: Genitalia, ventral. 85: Posterior portion of ninth tergite, dorsal. 86: Gonostylus, ventral. 87: Genitalia, ventral. 88: Fourth flagellomere, lateral. 89: Ditto, medial. Scales 0.05 mm. Arrows refer to characters described in the diagnoses.