Winnertzia rickebasta, Jaschhof & Jaschhof, 2020

Jaschhof, Mathias & Jaschhof, Catrin, 2020, Reevaluation of species richness in Winnertzia (Diptera, Cecidomyiidae, Winnertziinae), with descriptions of 37 new species from Sweden, Peru and Australia, Zootaxa 4829 (1), pp. 1-72 : 15-17

Winnertzia rickebasta sp. nov.

Figs 17–19

Diagnosis. A medium-sized, dark-brown Winnertzia , distinguished from other species of the discreta group by a set of genitalic characters, as follows ( Fig. 17 ). The big gonostylus has dense, moderately large microtrichia apically and medially, of which some are clustered to form a small tuft near the broadly rounded, almost inflated apex (↓ 5). Of the gonocoxites, the large, U-shaped emargination has an extensive unsclerotized area basally (↓ 6) and the dorsal apodemes are short (↓ 7). The broad-triangular tegmen has indistinct flaps without microtrichia. The anterior edge of the aedeagal bulge is well-marked as a transverse line (↓ 8). The posterior edge of the ninth tergite is markedly emarginated. Females and preimaginal stages of W. rickebasta are unknown.

Other male characters. Body size 2.4 mm. Head. Eye bridge 4–5 ommatidia long dorsally. Antenna longer than half body. Scape and pedicel same size, both concolorous with flagellum. 12 flagellomeres, translucent sensilla present on all flagellomeres. Fourth flagellomere: neck 0.7 times as long as node; node 1.8 times as long as broad; sensory hairs numerous; both the lateral and medial translucent sensilla long, filiform, linear to slightly bent ( Figs 18–19 ). Palpus shorter than head height, 4 setae-bearing segments; apical segment longest of all. Labella fully developed. Thorax. Pronotal setae 14. Anepimeral setae absent. Lateral mediotergal microtrichia slightly enlarged. Parascutellar area bright, sharply contoured. Wing slightly shorter than body, 2.5 times as long as broad. Costal cell reinforced. M 4 long, very slightly bent, extending to edge of wing. CuA moderately bent, declining before edge of wing. Legs. Scales pointed. Fore tibia 1.4 times length T 2. Acropods: claws slightly bent, 1 large and 1–2 smaller teeth basally; empodia vestigial. Abdomen. Pleural membrane setose. Genitalia ( Fig. 17 ). Ninth tergite longer than half gonopodal length; setae confined to posterior two thirds; anterior edge indistinct. Gonocoxal synsclerite broader than long; a short portion ventrobasally non-setose; ventroanterior edge slightly convex; ventro- and dorsoposterior portions ending at about same level. Gonostylus 2.3 times as long as broad, nearly parallel-sided; basolateral apophysis moderately large, angulated. Aedeagal apodeme parallel-sided, slightly constricted apically; solid basal portion short. Aedeagal bulge with broken rows of small spikes. Parameral apodemes moderately large.

Etymology. Rickebasta, used here as a noun in apposition, is a nature reserve in Uppland where the only specimen known of this species was collected.

Type material. Holotype. Male , Sweden, Uppland , Knivsta, Rickebasta Nature Reserve, swamp forest of alder interspersed with spruce trees, 12 June–24 July 2010, Malaise trap, M. & C. Jaschhof (spn GULI000021000 in NHRS).

Distribution and phenology. Winnertzia rickebasta is known from a single specimen, whose collection data are specified above.

Figures

FIGURES 14–19. Male morphology of Winnertzia pilosistylus (14–16) and Winnertzia rickebasta (17–19). 14: Fourth flagellomere, lateral, paratype. 15: Ditto, medial. 16: Genitalia, ventral, holotype. 17: Ditto. 18: Fourth flagellomere, lateral, holotype. 19: Ditto, medial. Scales 0.05 mm. Arrows refer to characters described in the diagnoses.