Falconina gracilis ( Keyserling, 1891 )

García, Fabián & Bonaldo, Alexandre B., 2023, Taxonomic revision of the soldier spider genus Falconina Brignoli, 1985 (Araneae: Corinnidae: Corinninae), Zootaxa 5343 (3), pp. 201-242 : 205-220

Falconina gracilis ( Keyserling, 1891)

Figs 1–9 , 25

Hypsinotus gracilis Keyserling, 1891: 56 , pl. 2, fig. 29 (♀ holotype from Rio Grande do Sul, Brazil, deposited in NHM 1890.7.1.1243, examined by photographs, Figs 9G–H ).

Corinna stigmatica Simon, 1897b (♀ holotype from Colonia Risso, Paraguay, deposited in MNHN, not examined, probably lost); synonymized with Falconina gracilis by Bonaldo, 2000: 79.

Corinna stigmatisata Simon, 1897a: 3 ; Simon, 1898a: 193, 196 (first revisor choice).

Corinna gracilis Petrunkevitch, 1911: 466 .

Corinna vertebrata Mello-Leitão, 1939 , figs 70–71 (♀ holotype from Capito Punto , Paraguay, Museum d’Histoire Naturelle Bale , MHNB, not re-examined); Mello-Leit ã o, 1940: 54, pl. 1, fig. 2; Mello-Leit ã o, 1941: 176, fig. 67; synonymized with Corinna stigmatica by Mello-Leit ã o, 1945: 222.

Corinna cribosa Mello-Leitão, 1942: 417 , fig. 42 (♀ holotype from Resistencia , Chaco, Argentina, XI.1939, M. Birabén leg., deposited in Museo de la Plata, MLP 15540 , not re-examined); synonymized with Falconina gracilis by Bonaldo, 2000: 79.

Falconina gracilis Bonaldo, 2000: 75 , figs 35–38, 41, 42, 63, 101, 215–228; Bonaldo & Brescovit, 2005; Valle et al., 2013: figs 1–3; Ramírez, 2014: 17; Wheeler et al., 2017: 588; Azevedo et al., 2022: 5 ( F. grassa , lapsus).

Diagnosis. Males of F. gracilis resemble those of F. taita sp. nov. by the presence of a short and thick apical spur on the RTA ventral lobe ( Fig. 5B ). They can be distinguished by the presence of a dorsal process of the ventral lobe of the RTA, a much smaller median lobe (robust in F. taita sp. nov.), and a dorsal lobe with multiple small, projected branches ( Bonaldo 2000: figs 224–225) (without apical spiniform projections in F. taita sp. nov.). Females resemble those of F. albomaculosa by the posterior margin of epigynal plate with a median notch ( Fig. 6B ), posterior vulvar plate slightly projected posteriorly, barely visible in ventral view and sub-squared in dorsal view, but can be differentiated by the margins of the epigynal plate and posterior vulvar plate divergent in ventral view (parallel in ventral view in F. albomaculosa ); dorsally, posterior vulvar plate with recurved anterior margin (anterior margin procurved in F. albomaculosa ) ( Fig. 6C ); secondary spermathecae inserted subapically in relation to primary spermathecae ( Fig. 6D ).

Description. See Bonaldo 2000: 79–83. Habitus of live specimens in Figs 3 and 7 .

New records. CUBA: Granma: El Mango, Rio Cauto [N20.55947°, W76.90953°, 12 m], M.A. Mendonça leg., 23.IV.2003, 1 ♀ ( IBSP 236874 ) . Guantánamo: Piedra La Vela, Camino al Rio Toa [N20.32277°, W74.86797°, 457 m], 12.XII.1997, 1 ♂ ( IBSP 236886 ) . Holguín: Moa , La Melba [N20.66158°, W74.95301°, 22 m], M.S. Da Silva leg., 21.IX.1997, 1 ♂ 3 ♀ ( IBSP 233839 ), J. Garlet et al. leg., 24.IX.1997, 1 ♂ ( IBSP 169764 ), M.S. Da Silva leg., same data, 1 ♀ ( IBSP 237079 ) . Santiago de Cuba: El Olimpo, La Gran Piedra [N20.00974°, W75.62721°, 711 m], 7.II.2000, 1 ♂ ( IBSP 236893 ) ; La Providencia [N20.00191°, W75.82708°, 37 m], A. Sánchez Ruiz leg., 30.IV.2004, 1 ♀ ( IBSP 236867 ) ; Pico CUBA [N19.94365°, W76.86111°, 24 m], R. Angrinah leg., 27.IX.1999, 1 ♂ 1 ♀ ( IBSP 233815 ) . PERÚ: Cusco: Rio Camisea, Cashiari [S11.82542°, W72.81947°, 421 m], J. Suarez & S. Cordova leg., V–VI.1997, 1 ♀ ( MUSM 0504313 ) . Madre de Dios: Zona reservada Tambopata [S12.55949°, W69.19081°, 193 m], C. Cañas leg., 1996, 1 ♀ ( MUSM 0503335 ) . BRAZIL: Amazonas: Benjamin Constant [S4.38421°, W70.0456°, 90 m], P.S. Pompeu et al. leg., 2014, 4 ♂ ( IBSP 238882 ) , 2 ♂ 1 ♀ ( IBSP 238980 ) , 2 ♂ ( IBSP 238909 ) , 1 ♀ ( IBSP 238790 ) , 2 ♂ ( IBSP 238825 ) , 1 ♂ 1 ♀ ( IBSP 238858 ) , 1 ♀ ( IBSP 238894 ) , 1 ♂ ( IBSP 238898 ) , 1 ♂ ( IBSP 238949 ) , 1 ♀ ( IBSP 238966 ) , 1 ♂ ( IBSP 238973 ) , 1 ♂ ( IBSP 238977 ) , 1 ♂ ( IBSP 239034 ) , 1 ♀ ( IBSP 239058 ) ; Manaus, Campus INPA [S3.09541°, W59.98856°, 49 m], T.R. Gasnier leg., II.1992, 1 ♂ ( MCN 22258 ) ; Colina de Aleio [S3.06547°, W59.96037°, 48 m], T.R. Gasnier leg., 1992, 1 ♂ ( MCN 22263 ) ; Manaus, Parque de 10 Novembro [S3.08527°, W60.00744°, 39 m], A.D. Brescovit leg., 10.I.1994, 1 ♂ ( MCN 25000 ) , H. Höfer leg., XI.1993 ( MCN 25001 ) , A.D. Brescovit leg., 5–10.I.1994, 1 ♂ ( MCN 24999 ) ; Urucu-Coari, Base de Operações Geólogo Pedro de Moura (S4.86878°, W65.26489°, 83m), A.B. Bonaldo, D. Candiani & M. Fernandes leg., 1.X.2004, 1 ♂ ( MPEG 010004 ) . Pará: Almeirim, Estação Ecológica (S0.8329°, W52.59434°, 89m), T. Gardner & M.A. Ribeiro Júnior leg., 7.III.2005, 1 ♂ ( MPEG 003769 ) , same collectors, 11.III.2005, 1 ♂ ( MPEG 37229 ) , T. Gardner leg., 7.IX.2004, 1 ♂ ( MPEG 003768 ) , T. Gardner & M.A. Ribeiro Júnior leg., 11.II.2005, 1 ♂ ( MPEG 003767 ) , T. Gardner leg., 7.IX.2004 ( MPEG 003771 ) ; Altamira , Castelo dos Sonhos [S8.31704°, W55.09853°, 253 m], M. Hoogmoed leg., 17.XI.2005, 1 ♀ ( MPEG 004515 ) ; Alter do Chão, Santarém [S2.50467°, W54.9548°, 35 m], H. Höfer leg., 25.I.1994, 1 ♂ ( MCN 25005 ) ; Belém, Campus Pesquisa MPEG [S1.45121°, W48.44536°, 16 m], E. Cafofo leg., 1.X.2010, 1 ♀ ( MPEG 37225 ) , A.B. Bonaldo leg., 17.IV.2009, 1 ♀ ( MPEG 37226 ) ; Campus UFPA [S1.4733°, W48.45727°, 6 m], J.A.P. Barreiros leg., 11.IX.2001, 1 ♂ ( MPEG 37227 ) ; Condomínio Alto de Pinteiros, Pratinha [S1.36603°, W48.47606°, 6 m], A.B. Bonaldo leg., 2.IV.2007, 1 ♂ ( MPEG 37228 ) ; Condomínio Jardim Universitário, Guamá [S1.47081°, W48.45583°, 6 m], A.B. Bonaldo leg., 3.III.2017, 1 ♂ ( MPEG 37230 ) , same collector, 4.I.2016, 1 ♂ ( MPEG 37231 ) ; Embrapa (S1.43584°, W48.44174°, 16 m), H.H.G. Oliveira leg., 22.XII.2014, 1 ♂ ( MPEG 031528 ) , same collector, 4.IX.2014, 1 ♀ ( MPEG 031527 ) ; Igarapé-Açu, Feiga UFRA [S1.4562°, W48.43769°, 44 m], 9.XII.2002, 1 ♂ ( MPEG 37232 ) ; Igarapé-Açu [S1.13033°, W47.62828°, 44 m], L.R. Fernandes leg., 23.II.2011, 3 ♂ 1 ♀ ( MPEG 37233 ) , same collector, 19.V.2011, 1 ♂ ( MPEG 37234 ) , same collector, 13.IX.2011, 1 ♂ ( MPEG 37235 ) , same collector, 19.V.2011, 2 ♂ ( MPEG 37224 ) , same collector, 14.II.2012, 1 ♂ ( MPEG 37236 ) ; Ilha de Marajó, Soure [S0.7345°, W48.5069°, 8 m], T.G. Pinheiro leg., 27–28.IX.2016, 1 ♀ ( IBSP 210848 ) ; Uruará [S3.72135°, W53.74343°, 139 m], T.G. Pinheiro leg., 2015, 1 ♀ ( IBSP 230886 ) , 1 ♀ ( IBSP 230936 ) ; Parauapebas , Canaã dos Carajás, [S6.53008°, W49.85177°], J. Lopes leg., 9.III.2005, 1 ♂ ( IBSP 51225 ) ; Canaã dos Carajás , Gruta NV 02 [S6.53008°, W49.85177°], I.C.F. Martins leg., 22–28.II.2005, 1 ♂ ( IBSP 55363 ) . Maranhão: Barreirinhas , Parque Nacional dos Lençóis Maranhenses (S2.71938°, W42.68682°, 4 m), J. Garlet et al. leg., 12– 18.X.2001, 1 ♂ ( IBSP 76687 ) ; Caxias, Universidade Estadual do Maranhão [S2.57724°, W44.20946°, 35 m], J. Garlet et al. leg., 2008, 1 ♀ ( IBSP 122434 ) ; Santa Quitéria do Maranhão, Fazenda Filadélfia [S3.51556°, W42.54667°, 26 m], J. Garlet et al. leg., 21.I.1999, 1 ♂ ( IBSP 36753 ) , J. Garlet et al. leg., 11.XII.1999, 6 ♂ ( IBSP 36083 ) . Ceará: Crato, Floresta Nacional do Araripe [S7.33839°, W39.41191°, 665 m], R. Azevedo leg., 22.IV2012, 1 ♂ ( MPEG 024479 ) , 1 ♂ ( MPEG 24480 ) ; Fortaleza, Itaperi [S3.79089°, W38.54782°, 29 m], M. Mourão leg., 8.VII.2011, 1 ♂ ( IBSP 164390 ) , Universidade Estadual do Ceará, Campus do Itaperi [S3.7878°, W38.55348°, 29 m], A. J. Santos leg., 30.VII.2007, 1 ♀ ( IBSP 97896 ) . Acre: Rio Branco [S9.94149°, W67.83331°, 158 m], R.S. Oliveira & B. Bezerra leg., 9.XI.1995, 1 ♂ ( IBSP 7013 ) , R.S. Vieira leg., na data, 1 ♂ ( IBSP 7018 ) . Piauí: Jaicós (S7.35889°, W41.13778°, 311 m), 2013–2015, 1 ♀ ( IBSP 231786 ) , T. Falótico leg., same data, 1 ♀ ( IBSP 231792 ) , 1 ♂ ( IBSP 231734 ) , O. Rocha leg., same data, 1 ♀ ( IBSP 231739 ) , A. Tonhasca Júnior leg., same data, 1 ♂ ( IBSP 231741 ) , 1 ♀ ( IBSP 231747 ) , A.M. Nunes leg., same data, 1 ♂ ( IBSP 231761 ) , E.F. Ramos leg., same data, 1 ♂ ( IBSP 231762 ) , C. Sampaio, F. Uemori & C.T. Olivares leg., same data, 1 ♂ ( IBSP 231768 ) , Equipe Biota leg., same data, 1 ♂ ( IBSP 231783 ) , 1 ♂ ( IBSP 231784 ) , 1 ♂ ( IBSP 231785 ) , Equipe UFRJ leg., same data, 1 ♀ ( IBSP 231787 ) , Equipe Biota leg., same data, 1 ♀ ( IBSP 231793 ) ; Parnaíba, Distrito de Irrigação dos tabuleiros Litorâneos do Piauí ( Ditalpi ) [S3.03075°, W41.7717°, 39 m], M. Maia leg., 2008, 1 ♀ ( IBSP 143979 ) ; Parnaíba , Campus Embrapa Meio Norte [S3.08607°, W41.7826°, 53 m], M. Maia leg., 2007, 1 ♂ ( IBSP 86570 ) ; São João do Piauí, Parque Nacional da Serra da Capivara (S8.41667°, W42.33333°, 305 m), M. Ohlweider leg., IX.2007 – X.2010, 1 ♂ 1 ♀ ( IBSP 225212 ) , K.C. Ohweiler leg., same data, 1 ♂ ( IBSP 225224 ) ; Teresina , Campus Embrapa Meio Norte [S5.0372°, W42.79938°, 66 m], R.G. Bussel & L. Indrusiak leg., 1.VI.2005, 1 ♂ ( IBSP 55963 ) . Tocantins: Araguaína, Setor Cimba [S7.1874°, W48.20134°, 253 m], S. Barros leg., 15.VII.2014, 1 ♀ ( IBSP 237263 ) ; UFT-Cimba [S7.18229°, W48.19738°, 253 m], V.E. Sandoval leg., 11.V.2017, 1 ♀ ( IBSP 237282 ) ; Palmas, Entrada da Fazenda Brejo Redondo [S10.2314°, W48.29352°, 276 m], I. Knysak; R. Martins leg., 11–19.VI.2001, 1 ♀ ( IBSP 214777 ) ; Porto Nacional [S10.70115°, W48.40637°, 243 m], R. Martins & G. Puorto leg., 18–23.III.2001, 1 ♀ ( IBSP 219495 ) , I. Knysak & R. Martins leg., 14–23.II.2003, 1 ♀ ( IBSP 224693 ) , 1 ♂ ( IBSP 224697 ) , 1 ♂ ( IBSP 224706 ) , 1 ♀ 1 imm. ( IBSP 224713 ) . Mato Grosso: Alta Floresta, Fazenda Anacã [S9.84799°, W56.10205°, 272 m], J. Garlet et al. leg., 25.VII.2016 – 27.XI.2017, 3 ♂ 1 ♀ ( IBSP 232893 ) , same collector, 2017, 1 ♂ ( IBSP 232096 ) , same collector, 25.VII.2016 – 27.XI.2017, 2 ♂ ( IBSP 232897 ) , same collector, 2017, 1 ♀ ( IBSP 232053 ) , 1 ♂ ( IBSP 232054 ) , M.S. Da Silva leg., same data, 7 ♂ ( IBSP 232055 ) , 1 ♀ ( IBSP 232068 ) , 1 ♂ ( IBSP 232079 ) , 2 ♂ 1 ♀ ( IBSP 232084 ) , 1 ♂ ( IBSP 232092 ) , 1 ♀ ( IBSP 232093 ) , 1 ♀ ( IBSP 232103 ) , same collector, 25.VII.2016 – 27.XI.2017, 1 ♂ ( IBSP 232888 ) , N.M. Rocha leg., same data, 1 ♂ ( IBSP 232889 ) , 1 ♂ ( IBSP 232895 ) , 1 ♂ ( IBSP 232901 ) , 1 ♂ ( IBSP 232902 ) , 2 ♀ ( IBSP 232903 ) , R.A.K. Ribeiro leg., same data, 1 ♀ ( IBSP 232913 ) , 1 ♀ ( IBSP 232923 ) ; Cáceres, Bairro Centro [S16.07106°, W57.68703°, 147 m], S.K.A. Barros leg., 21.XI.2004, 1 ♂ ( IBSP 70887 ) , D. Batistella leg., 20.XI.2004, 1 ♀ ( IBSP 70902 ) , Bairro Jardim Guanabara [S16.09293°, W57.66561°, 147 m], S.K.A. Barros leg., 19.XI.2004, 1 ♂ ( IBSP 70883 ) , D. Batistella leg., same data, 1 ♂ ( IBSP 70898 ) , same collector, 17.VIII.2004, 1 ♀ ( IBSP 70905 ) ; Cuiabá, Bairro Santa Cruz [S15.60841°, W56.04536°, 185 m], D. Batistella leg., 12.VI.2005 – 26.IV.2006, 1 ♂ ( IBSP 63180 ) ; Nova Mutum (S13.82889°, W56.08222°, 436 m), D. Batistella leg., 6–15.VIII.2001, 2 ♀ ( IBSP 29537 ) ; Poconé, Fazenda Santa Ines [S16.25025°, W56.64586°, 173 m], A. Lise & A. Braul Jr leg., 3.VII.1992, 1 ♂ ( MCP 2423 ) , Same locality [S16.26809°, W56.63202°, 173 m], D. Batistella leg., 2010–2012, 2 ♂ 2 ♀ ( IBSP 249219 ) , 1 ♀ ( IBSP 249206 ) , 1 ♀ ( IBSP 249212 ) , 1 ♂ 1 ♀ ( IBSP 249223 ) , 1 ♀ ( IBSP 249201 ) , 1 ♀ ( IBSP 249202 ) , 1 ♀ ( IBSP 249203 ) , 1 ♀ ( IBSP 249204 ) , 1 ♀ ( IBSP 249205 ) , 1 ♂ ( IBSP 249207 ) , 1 ♂ ( IBSP 249208 ) , 2 ♀ ( IBSP 249209 ) , 1 ♂ ( IBSP 249210 ) , 1 ♀ ( IBSP 249211 ) , 1 ♀ ( IBSP 249213 ) , 1 ♀ ( IBSP 249214 ) , 1 ♀ ( IBSP 249215 ) , Equipe IBSP leg., same data, 1 ♀ ( IBSP 249216 ) , 3 ♂ 2 ♀ ( IBSP 249217 ) , 1 ♀ ( IBSP 249218 ) , 3 ♀ ( IBSP 249220 ) , 1 ♂ 4 ♀ ( IBSP 249221 ) , D.F. Candiani & C.A.R. Souza leg., same data, 1 ♀ ( IBSP 249222 ) , J.P. Guadanucci & C. Fukushima leg., same data, 1 ♀ ( IBSP 249224 ) ; Santo Antonio de Levergere [S15.86417°, W56.07917°, 155 m], H. Duarte leg., 30.IV.1982, 1 ♀ ( MCP 2533 ) , same collector, 9.X.1982, 1 ♂ ( MCP 2537 ) , same collector, 13.X.1982, 1 ♂ ( MCP 2531 ) , same collector, 9.X.1982, 1 ♂ ( MCP 2534 ) ; Sinop, Embrapa Agrossilvipastoril [S11.87189°, W55.59712°, 368 m], D. Batistella leg., 23.II.2015, 1 ♂ ( IBSP 168004 ) [S11.85273°, W55.51532°, 373 m], D. Batistella leg., 6.I.2014, 1 ♂ ( IBSP 170111 ) . Rondônia: Ouro Preto do Oeste, Campo Experimental da Embrapa [S10.73152°, W62.25617°, 245 m], L.S.P. Trigueiro et al., leg., VIII.2008 – IX.2008, 1 ♀ ( IBSP 124603 ) , same collector, VIII.2008 – IX.2008, 1 ♂ ( IBSP 124695 ) , 1 ♀ ( IBSP 124692 ) , VIII.2008 – IX.2008, 1 ♀ ( IBSP 124597 ) , 1 ♀ ( IBSP 124598 ) , 1 ♂ ( IBSP 124599 ) , 1 ♂ ( IBSP 124600 ) , 1 ♂ ( IBSP 124601 ) , 1 ♀ ( IBSP 124602 ) , 1 ♀ ( IBSP 124691 ) , M. Carvalho leg., same data, 1 ♂ ( IBSP 124693 ) , C. Carlos et al., leg., same data, 1 ♀ ( IBSP 124694 ) , J.M. M. leg., same data, 1 ♀ ( IBSP 124696 ) , 1 ♀ ( IBSP 124697 ) , J. Barp leg., same data, 1 ♀ ( IBSP 124698 ) , V.C. Onófrio leg., same data, 1 ♂ ( IBSP 124699 ) , 1 ♀ ( IBSP 124700 ) , V.C. Onofrio leg., same data, 1 ♂ ( IBSP 124701 ) , L. Marconini leg., same data, 1 ♀ ( IBSP 124702 ) , D.P. Pinseta leg., same data, 1 ♂ ( IBSP 124703 ) , Equipe IBSP leg., same data, 1 ♂ ( IBSP 124704 ) , R. Camargo leg., same data, 1 ♀ ( IBSP 124705 ) , R.L. Lima leg., same data, 1 ♀ ( IBSP 124706 ) , same data, 1 ♀ ( IBSP 124707 ) , A.J. Santos leg., same data, 1 ♀ ( IBSP 124708 ) ; Vilhena [S12.75239°, W60.14258°, 584 m], A.J. Santos leg., IX.1999, 1 ♂ ( IBSP 80977 ) . Rio Grande do Norte: João Câmara (S5.5375°, W35.81972°, 157m), N. Sebastian et al. leg., VII.2012, 2 ♂ ( MPEG 019973 ) , same collector, VI.2013, 1 ♀ ( MPEG 019976 ) , same collector, VII.2013, 1 ♂ ( MPEG 019975 ) , same collector, IX.2013, 1 ♀ ( MPEG 019974 ) , same collector, XI.2013, 1 ♂ ( MPEG 019971 ) , same collector, VII.2012, 1 ♂ ( MPEG 019972 ) ; Serra Negra do Norte, Estação Ecológica do Seridó [S6.58058°, W37.28324°, 219 m], L.S.P. Trigueiro, C.A.D. Teixeira, L.A.P. Sallet & K.S. Serra leg., VI.2008, 1 ♂ ( IBSP 123402 ) , same collectors leg., 10–21.XII.2007, 1 ♂ ( IBSP 123454 ) . Paraíba: São João do Cariri [S7.38898°, W36.5331°, 458 m], T.G. Pinheiro leg., X.2003, 1 ♀ 1 imm. ( IBSP 50858 ) ; São José dos Cordeiros, RPPN Fazenda das Almas [S7.47917°, W36.905°, 667 m], T.G. Pinheiro leg., XII.2006 – IX.2009, 1 ♂ ( IBSP 220294 ) , 1 ♂ ( IBSP 220300 ) ; field number PEMPF 0056, J.B.R Alencar leg., 5.VIII.2011, 1 ♀ (CHNUFPI 2439). Pernambuco: Brejão , IPA [S9.03068°, W36.56645°, 778 m], J.A.S. Cunha leg., 27.VI.2012, 2 ♀ ( IBSP 232538 ) . Sergipe: Canindé de São Francisco, Usina Hidrelétrica de Xingó [S9.62258°, W37.78786°, 203 m], 16.XI.2000, 1 ♂ ( IBSP 88319 ) , same collector, 4.III.2000 – 29.III.2001, 2 ♂ ( IBSP 88320 ) , same collector, 11.X.2001, 1 ♂ ( IBSP 88335 ) , same collector, 4.III.2000 – 29.III.2001, 2 ♂ ( IBSP 88348 ) . Bahia: Catu [S12.35611°, W38.37401°, 120 m], M.F. Dias leg., 10.X.1998, 1 ♀ ( IBSP 40652 ) ; Central, Toca dos Índios, F.S. Cunha & E.F. Ramos leg., III.2002, 1 ♂ ( IBSP 247549 ) ; Toca dos Pilões, F.S. Cunha & E.F. Ramos leg., 5–12.III.2002, 1 ♂ ( IBSP 247531 ) [S11.13524°, W42.11046°, 636 m], E.F. Ramos leg., 16.VII.1997, 3 ♂ 1 ♀ ( IBSP 13368 ) , same locality (S11.31667°, W42.16667°, 934 m), E.F. Ramos & S.F. de Paula leg., 27.II–15.III.2002, 1 ♂ ( IBSP 222822 ) , 1 ♀ ( IBSP 222824 ) , same locality [S11.13524°, W42.11046°, 698 m], F.S. Cunha & E.F. Ramos leg., 2–8.III.2003, 3 ♂ 2 ♀ ( IBSP 247078 ) ; Curaçá, Riacho Seco [S8.7767°, W39.67077°, 346 m], I. Quinet & R. Azevedo leg., VI.2010, 1 ♂ ( IBSP 249040 ) , R. Azevedo leg., same data, 1 ♀ ( IBSP 249039 ) ; Ilhéus, Olivença [S14.94763°, W39.02044°, 43 m], M.F. Dias leg., VIII.1998, 1 ♀ ( IBSP 35914 ) ; Jandaíra [S11.56734°, W37.78364°, 109 m], T. Silva leg., XII.2010, 1 ♂ ( IBSP 170143 ) ; Jussara, Toca da Esperança [S11.051°, W41.97298°, 636 m], E.F. Ramos & F.S. Cunha leg., 18–25.VII.2002, 1 ♀ ( IBSP 238250 ) ; Lauro de Freitas [S12.88676°, W38.29821°, 14 m], H.F. Fontana leg., 7.III.2014 – 10.VII.2014, 2 ♂ 1 ♀ ( IBSP 169327 ) ; Maracás, Porto Alegre, Barragem da Pedras Altas [S11.18699°, W40.05313°, 367 m], L.P. de Oliveira leg., 6.XII.2006, 1 ♂ ( IBSP 80890 ) ; Mato de São João, Imbasai [S12.48035°, W37.96403°, 43 m], M. Dias leg., 29–10.III.2008, 1 ♂ ( IBSP 118796 ) ; Miguel Calmon , Parque Estadual das Sete Passagens [S11.3922°, W40.54012°, 830 m], B.R.S. Pereira leg., 2017, 1 ♂ 1 ♀ ( IBSP 227462 ) ; Milagres [S12.86687°, W39.86124°, 377 m], V.E.F. Santana et al., leg., 8.I.2010, 1 ♂ ( IBSP 238031 ) , same collector, 28.I.2010, 1 ♀ ( IBSP 238079 ) ; Salvador, Baia do Aratu [S12.77381°, W38.49851°, 23 m], K. Benati leg., 2008, 1 ♀ ( IBSP 145853 ) , 1 ♂ ( IBSP 145856 ) , same locality (S12.78333°, W38.46667°, 23 m), K. Benati & S. Vareja leg., same data, 1 ♂ ( IBSP 145753 ) ; Ondina [S13.00581°, W38.50918°, 0 m], 2013, 1 ♀ ( IBSP 210727 ) ; Parque Municipal de Pituaçu [S12.96806°, W38.41103°, 0 m], E. Alves leg., 27–31.V.2004, 1 ♂ ( IBSP 120353 ) , 1 ♂ ( IBSP 120361 ) , 1 ♂ ( IBSP 120365 ) , 1 ♂ ( IBSP 120366 ) , 1 ♀ ( IBSP 120368 ) , same locality [S12.96423°, W38.42308°, 18 m], same collector and date, 1 ♀ ( IBSP 120354 ) ; Parque Municipal do Pituaçu [S12.96806°, W38.41103°, 0 m], E. Alves leg., 27–31.V.2004, 1 ♀ ( IBSP 80729 ) , 1 ♀ ( IBSP 80793 ) , 1 ♀ ( IBSP 80798 ) , Parque Zoobotânico de Salvador [S13.00926°, W38.50483°, 0 m], 2013, 2 ♀ ( IBSP 210616 ) , 1 ♀ ( IBSP 210618 ) , 1 ♂ 1 ♀ ( IBSP 210623 ) , 1 ♂ ( IBSP 210633 ) ; Una, Reservatório Ecológico do Una (S15.29667°, W39.07444°, 31 m), M.F. Dias leg., X.1999 – IX.2000, 1 ♂ ( IBSP 64555 ) . Goiás: Caldas Novas, Parque Estadual Caldas Novas [S17.7701°, W48.65652°, 735 m], A. Anker & F. García leg., 31.I.2021, 1 ♂ ( MPEG 37220 ) ; Catalão (S18.17°, W47.94194°, 814 m), L.C. Bergamini leg., 2002, 1 ♂ ( IBSP 38855 ) , C.A. Rheims leg., same data, 1 ♂ 1 ♀ ( IBSP 38856 ) ; Goiânia, Campus Samambaia UFG, Instituto de Química (S16.60333°, W49.26194°, 746 m), A. Anker & F. García leg., 7.VII.2020, 1 ♀ ( MPEG 37221 ) , same collector, 15.VII.2020, 1 ♂ 2 ♀ ( MPEG 37222 ) ; Campus Samambaia UFG, Instituto IV Ciências Biológicas (S16.60163°, W49.26014°, 746 m), A. Anker & F. García leg., 10.VII.2020, 5 ♀ ( MPEG 37223 ) [S16.68341°, W49.27349°, 762 m], C.A. Rheims leg., 2008, 2 ♀ ( IBSP 122608 ) ; Mineiros , Parque Nacional das Emas [S17.92001°, W52.8072°, 759 m], J.M. Bortolin leg., 22.IV.2000, 1 ♀ ( IBSP 26272 ) , same collector, 22.IV.2001, 1 ♀ ( IBSP 26274 ) ; Montividiu [S17.4525°, W51.17898°, 846 m], Equipe Biota leg., 2008, 1 ♂ 1 imm. ( IBSP 145379 ) ; same locality [S17.44727°, W51.17621°, 846 m], F. Limeira de Oliveira leg., same data, 1 ♀ ( IBSP 145380 ) . Distrito Federal: Brasilia, EPPN Trecho 01, Chácara 97, Lago Norte [S15.73872°, W47.85937°, 1033 m], R. Góes leg., 7.III.2003, 1 ♀ ( UNB2412 ) ; Brasília, Parque Way [S15.89034°, W47.9622°, 1087 m], T. Souza et al., leg., 26.IV.2006, 1 ♀ ( IBSP 62711 ) . Espírito Santo: Cariacica, Alto do Alegre, Reserva Estadual de Duas Bocas (S20.27294°, W40.47667°, 241 m), J.P.P. Pena-Babosa et al., leg., 13–14.X.2005, 1 ♀ ( IBSP 133200 ) ; Conceição da Barra, Floresta Nacional do Rio Preto, Trilha do Mata-Pau (S18.38773°, W39.85091°, 53 m), P.C. Motta leg., 19.X.2005, 4 ♂ 3 ♀ ( IBSP 135109 ) ; Linhares [S19.40002°, W40.07103°, 25 m], P.C. Motta leg., 9–15.I.2012, 1 ♂ ( IBSP 249641 ) . Minas Gerais: Araguari, Bosque John Kennedy (S18.65158°, W48.18264°, 948 m), E.S.S. Alvares, C.S. Azevedo & F.O. Machado leg., I–XII.2010, 1 ♀ ( IBSP 226194 ) , 1 ♂ ( IBSP 226261 ) , R. Gallon leg., same data, 1 ♂ ( IBSP 226264 ) ; Araguari, Uberlândia, Usina Hidrelétrica de Capim Branco [S18.77649°, W48.1639°, 758 m], A.J. Santos leg., 2.II.2005, 1 ♀ ( IBSP 71129 ) ; Belo Horizonte, Campus UFMG, Estaç ã o Ecológica [S19.87373°, W43.97241°, 815 m], M.E. Bichuette leg., 17.VI.1994, 1 ♀ ( IBSP 26933 ) , E.S.S. Álvares leg., 27.III.2000, 3 ♂ 7 ♀ ( IBSP 27236 ) , L. Almeida Silva leg., same data, 1 ♂ 1 ♀ ( IBSP 27237 ) , J.P.R. Brichta leg., same data, 1 ♂ 1 ♀ ( IBSP 27239 ) ; Estação Ecológica da UFMG, J.P.R. Brichta leg., IX.1999 – II.2001, 30 ♂ 7 ♀ ( IBSP 32294 ) , 10 ♂ 6 ♀ ( IBSP 32324 ) , same collector, VII.1999 – II.2001, 3 ♂ 7 ♀ ( IBSP 32484 ) , M.A. Benamu leg., same data, 10 ♂ 5 ♀ ( IBSP 32509 ) , E.S.S Álvares, E.O. Machado & C.S. Azevedo leg., VII.1999, 6 ♀ ( IBSP 32268 ) ; Coromandel , Vale Rio Santo Inácio [S18.48417°, W47.20649°, 934 m], M.A. Benamu leg., 12.V.2001, 1 ♀ ( IBSP 28567 ) ; Juatuba, Área de Preservação de Serra Azul [S19.95194°, W44.34278°, 764 m], M.A. Benamu leg., IV.2002, 1 ♂ ( IBSP 35841 ) ; Pains [S20.37544°, W45.65186°, 753 m], Andrade & Arnoni leg., 3.XI.2005, 1 ♂ ( IBSP 71895 ) ; Prudente de Morais, Fazenda Sapé [S19.48039°, W44.1511°), R.L. Caraíba leg., II.2003, 1 ♂ ( IBSP 44208 ) ; Rio Acima, Estação Biológica Estadual do Panga [S19.18199°, W48.3947°], V. F. Mattos leg., II–VI.2018, 1 ♀ ( IBSP 247358 ) ; Próximo da Serra da Gandarela [S19.18199°, W48.3947°], R.L. Caraíba leg., 5–6.IX.2009, 2 ♀ ( IBSP 149289 ) ; Uberlândia, Estação Biológica Estadual do Panga [S19.18199°, W48.3947°], V. F. Mattos leg., II–VI.2018, 1 ♀ ( IBSP 247359 ) , same locality [S19.18199°, W48.3947°], E.C. Iablanski leg., same data, 1 ♀ ( IBSP 247360 ) , same locality [S19.18199°, W48.3947°], E.V.J. Nascimento leg., same data, 1 ♀ ( IBSP 247361 ) . Mato Grosso do Sul: Anaurilândia, Usina Hidrelétrica Engenheiro Sérgio Motta (S22.38306°, W52.81178°, 252 m), R. Bertani & C.F.Y. Fukami leg., 15.XI– 23.XII.1999, 1 ♂ 1 ♀ ( IBSP 23400 ) , same collector, 17–24.VII.2000, 1 ♂ 1 ♀ ( IBSP 30888 ) , F.S. Cunha & J.P.L. Guadanucci leg., 2000, 1 ♂ 1 ♀ ( IBSP 30964 ) , Inone et al., leg., 12–19.III.2001, 1 ♂ ( IBSP 38897 ) ; Usina Hidrelétrica Sérgio Motta, F.S. Cunha & C.A.R. Souza leg., 2000, 1 ♂ ( IBSP 31205 ) , F.S.Cunha & J.P. Guadanucci leg., 17– 24.VII.2000, 1 ♀ ( IBSP 31226 ) , D.F. Candiani & C.A.R. Souza leg., 2000, 1 ♀ ( IBSP 35239 ) , J.P. Guadanucci & C. Fukushima leg., 15–19.VIII.2000, 1 ♂ ( IBSP 35403 ) , K.O. Vieira leg., 8–10.VIII.2000, 1 ♀ ( IBSP 35707 ) , same collector, 9.VIII.2000, 2 ♂ ( IBSP 35735 ) ; Anaurilândia [S22.18521°, W52.72001°, 308 m], Inone et al., leg., 12– 19.III.2001, 6 ♂ 8 ♀ ( IBSP 41260 ) , same collector, 5–11.III.2001, 1 ♂ ( IBSP 41316 ) ; Bataguassu, Fazenda Orelha de Onça [S21.72368°, W52.43777°, 334 m], Inone et al., leg., 3.VIII.2000, 1 ♀ ( IBSP 35427 ) ; Bonito, Pitangueiras [S20.50656°, W54.6168°, 549 m], E.S.S. Alvares, C.S. Azevedo & F.O. Machado leg., X.2002, 1 ♂ ( IBSP 247617 ) ; Brazilândia, Usina Hidrelétrica Engenheiro Sérgio Motta, Fazenda Flórida [S22.50128°, W53.01345°, 240 m], Inone et al., leg., 2000, 1 ♂ 1 ♀ ( IBSP 35260 ) ; Camapuã, Fazenda Maravilha [S19.55661°, W54.04025°, 477 m], Inone et al., leg., 13.X.1998, 1 ♂ ( IBSP 37449 ) , same collector, 19.XII.1998, 1 ♂ 1 ♀ ( IBSP 37453 ) ; Campo Grande , RPPN.UFMS [S20.50656°, W54.6168°, 549 m], Inone et al., leg., 24.I.2014, 1 ♂ ( IBSP226626 ) , same collector, 13–27.I.2014, 1 ♀ ( IBSP 226623 ) , 1 ♂ ( IBSP 226624 ) , J. Raizer et al., leg., 18.I.2014, 1 ♂ ( IBSP 226627 ) , J. Raizer leg., 19.I.2014, 1 ♂ ( IBSP 226629 ) , A.J. Santos leg., 13.I.2014, 1 ♂ ( IBSP 226632 ) , R. Bessi leg., 15. I.2014, 1 ♂ ( IBSP 226633 ) , K. Anjos leg., 22.I.2014, 1 ♂ ( IBSP 226635 ) , same collector, 24.I.2014, 1 ♀ ( IBSP 226636 ) , same collector, 13.I.2014, 1 ♀ ( IBSP 226637 ) ; Corumbá, Passo da Lontra [S19.58026°, W57.02131°, 91 m], K. Anjos leg., IV.1998, 1 ♂ ( IBSP 21892 ) , same collector, IX.1999, 1 ♂ 2 ♀ ( IBSP 35051 ) ; Passo do Lontra , K. Anjos leg., IV.1999, 1 ♂ ( IBSP 33136 ) ; Dois Irmãos do Buriti, Piraputanga [S20.45577°, W55.50052°, 448 m], K. Anjos leg., 16–26.II.2008, 1 ♀ ( IBSP 128786 ) ; Dois Irmãos do Buriti , K. Anjos leg., 31.VI–6.VII.1999, 1 ♂ ( IBSP 222882 ) , 1 ♂ ( IBSP 222883 ) , 1 ♀ ( IBSP 222884 ) ; Dourados [S22.24911°, W54.81925°, 430 m], K. Anjos leg., VI.2004, 1 ♂ 1 ♀ ( IBSP 125060 ) , 2 ♂ ( IBSP 125054 ) , 2 ♂ 1 ♀ ( IBSP 125055 ) , 1 ♂ 1 ♀ ( IBSP 125056 ) , 1 ♀ ( IBSP 125057 ) , D. Araujo et al., leg., same data, 1 ♂ ( IBSP 125058 ) , 1 ♂ ( IBSP 125059 ) , 1 ♂ ( IBSP 125061 ) , 1 ♂ ( IBSP 125062 ) , 1 ♀ ( IBSP 125063 ) , 1 ♂ ( IBSP 125064 ) , R.R. Silva leg., same data, 2 ♂ ( IBSP 125065 ) , C.A. Rheims leg., same data, 1 ♀ ( IBSP 125066 ) , V.S. Moreira leg., same data, 1 ♂ ( IBSP 125067 ) ; Ivinhema , REA [S22.30772°, W53.82721°, 364 m], V.S. Moreira leg., VI.2004, 1 ♀ ( IBSP 125053 ) ; Mundo Novo, Bairro Berneck [S23.92926°, W54.28764°, 298 m], S.T.M.Guerra leg., IV–VI.2008, 1 ♂ 4 ♀ ( IBSP 143675 ) ; Bairro Centro , V.S. Moreira leg., IV–VI.2008, 1 ♂ ( IBSP 143596 ) ; Bairro Copagril , E.S.S. Alvares, C.S. Azevedo & F.O. Machado leg., IV–VI.2008, 2 ♂ 2 ♀ ( IBSP 143713 ) , 1 ♂ 1 ♀ ( IBSP 143712 ) ; Bairro Tapajós , R. Mataveli leg., IV–VI.2008, 1 ♂ ( IBSP 143627 ) , E.S.S. Alvares, C.S. Azevedo & F.O. Machado leg., same data, 2 ♂ ( IBSP 143628 ) ; Paranaíba [S19.67722°, W51.19083°, 395 m], E.S.S. Alvares, C.S. Azevedo & F.O. Machado leg., VIII.1983, 1 ♂ ( IBSP 6775 ) . Rio de Janeiro: Campos dos Goytacazes [S21.76679°, W41.3325°, 10 m], L. Indrusiak leg., 1995, 1 ♂ 1 ♀ ( IBSP 6925 ) , R. Camargo leg., same data, 1 ♀ ( IBSP 6926 ) ; Mendes (S22.52667°, W43.73278°, 479 m), R. Lampert leg., X.2003, 1 ♂ ( IBSP 40948 ) ; Nova Iguaçu, Reserva Biológica do Tinguá [S22.5804°, W43.39972°, 399 m], T. Gomes leg., II.2002, 1 ♂ ( IBSP 118087 ) ; RPPN dos Petroleiros [S22.59399°, W43.43054°, 139 m], L.S.P. Trigueiro, C.A.D. Teixeira, L.A.P. Sallet & K.S. Serra leg., 4–6.IV.2012, 2 ♀ ( IBSP 215342 ) ; Paraíba do Sul, Fazenda Maravilha (S22.2°, W43.26667°, 352m), L.S.P. Trigueiro et al., leg., 16–18.VIII.2001, 1 ♀ ( IBSP 221829 ) , 1 ♀ ( IBSP 221831 ) , L.S.P. Trigueiro, C.A.D. Teixeira, L.A.P. Sallet & K.S. Serra leg., same data, 1 ♂ ( IBSP 221832 ) ; Rio de Janeiro, Restinga da Marambaia (S23.06431°, W43.80225°, 0 m), L.S.P. Trigueiro, C.A.D. Teixeira, L.A.P. Sallet & K.S. Serra leg., 2013, 1 ♀ ( IBSP 213493 ) ; Volta Redonda, Floresta da Cicuta [S22.54153°, W44.08843°, 394 m], L.S.P. Trigueiro, C.A.D. Teixeira, L.A.P. Sallet & K.S. Serra leg., 11–18.VI.2001, 1 ♀ ( IBSP 223082 ) . São Paulo: Águas da Prata [S21.93667°, W46.71667°, 868 m], C. Bertim leg., 10–15.I.1999, 1 ♂ ( IBSP 20784 ) , Equipe Biota leg., 29.XII.2000, 1 ♀ ( IBSP 27788 ) , same locality [S21.93759°, W46.72068°, 868 m], Y. Pesek leg., IX.2004, 3 ♂ 1 ♀ ( IBSP 70981 ) ; Araraquara [S21.78341°, W48.19499°, 665 m], A. Girotti et al., leg., 2003, 1 ♂ ( IBSP 43015 ) ; Atibaia (S23.11694°, W46.55028°, 777m), 28.X.1998, 2 ♂ ( IBSP 20330 ) ; Bernardino de Campos [S23.01555°, W49.47092°, 672 m], 28.V.2004, 1 ♀ ( IBSP 44299 ) , same locality (S23.01306°, W49.47417°, 672 m), A.C.B. Silveira leg., 5.XI.2003, 1 ♀ ( IBSP 40941 ) ; Biritiba Mirim, Barragem do Rio Biritiba [S23.60376°, W46.08628°, 780 m], V.2003, 2 ♀ ( IBSP 122419 ) ; Botucatu [S22.85962°, W48.45528°, 794 m], A. Girotti leg., XI.2013, 3 ♂ 1 imm. ( IBSP 215513 ) ; Caieiras [S23.36191°, W46.74789°, 775 m], S.H. Seike leg., 1.XI.2005, 1 ♂ ( IBSP 59538 ) ; Campinas, Campus Unicamp [S22.81853°, W47.06512°, 608 m], A.E.C. Farinha leg., No data, 1 ♂ ( IBSP 36362 ) (S22.90556°, W47.06083°, 608 m), A.E.C. Farinha leg., 16.VI.2003, 1 ♀ ( IBSP 39844 ) , same locality [S22.90582°, W47.07509°, 608 m], same collector, 14.II.2006, 1 ♂ ( IBSP 72975 ) , same locality (S22.90556°, W47.06083°, 683 m), A.J. Santos leg., 2.VIII.2003, 1 ♂ ( IBSP 40616 ) ; Catanduva, Usina São Domingos [S21.1007°, W49.01489°, 516 m], A.E.C. Farinha leg., 16.IX.2001, 1 ♀ ( IBSP 31894 ) ; Cotia [S23.6091°, W46.91886°, 865 m], J.A. de Almeida leg., 15.V.2008, 1 ♀ ( IBSP 116677 ) ; Dois Córregos, Sítio Guedes [S22.36792°, W48.37104°, 703 m], C. Bertim & M. Costa leg., 1–6.IV.2002, 1 ♀ ( IBSP 34968 ) ; Embu Guaçu (S23.83222°, W46.81139°, 762 m), O.A. Norales leg., IV.1976, 1 ♂ ( IBSP 2784 ) ; Francisco Morato [S23.2827°, W46.73988°, 834 m], T. Godói leg., 28.VII.2005, 1 ♂ ( IBSP 56992 ) ; Indaiatuba [S23.13147°, W47.23776°, 573 m], J.P. Guadanucci & R. Bertani leg., 7.I.2008, 1 ♀ ( IBSP 115462 ) ; Itirapina, Estação Ecológica de Itirapina [S22.24962°, W47.82735°, 813 m], Equipe IBSP leg., 2001, 1 ♀ ( IBSP 125423 ) , same collector, 14–21.V.2001, 1 ♀ ( IBSP 138433 ) ; Estação Ecológica Itirapina , D.F. Candiani & R.P. Indicatti leg., 14–21.V.2001, 1 ♂ 1 ♀ ( IBSP 222968 ) ; Itu, Camping Lara, Camping Casarão [S23.2683°, W47.22612°, 699 m], D. Araújo & A. Girotti leg., 2015, 1 ♀ ( IBSP 237110 ) [S23.26993°, W47.29042°, 592 m], D. Araújo & A. Girotti leg., 17.VII.2006, 1 ♂ ( IBSP 75807 ) , same locality (S23.26417°, W47.29917°, 579 m), R. D’Avilla leg., 20.I.2003, 1 ♂ ( IBSP 37204 ) ; Jaboticabal [S21.26509°, W48.32656°, 608 m], D. Araújo & A. Girotti leg., III – IV.2004, 3 ♂ 1 ♀ ( IBSP 55885 ) , 12 ♂ 6 ♀ ( IBSP 55884 ) , A. Girotti leg., same data, 2 ♂ 1 ♀ 1 imm. ( IBSP 55876 ) ; Jaú [S22.30807°, W48.56677°, 608 m], A. Girotti leg., 12.IV.2009, 1 ♀ ( IBSP 135301 ) ; Jundiaí (S23.18639°, W46.88417°, 733 m), A. Girotti leg., 11.X.1993, 1 ♂ 1 ♀ ( IBSP 26758 ) ; Louveira (S23.08639°, W46.95056°, 733 m), Alunos Curso UNESP leg., 10.X.1995, 1 ♂ 1 ♀ ( IBSP 12045 ) , T.G. Pinheiro leg., same data, 5 ♂ 3 ♀ ( IBSP 12047 ) , M.A. Rezende leg., 16.I.1996, 1 ♂ 1 ♀ ( IBSP 12063 ) , R. Pinto da Rocha leg., same data, 1 ♀ ( IBSP 12065 ) ; Martinópolis [S22.14953°, W51.17252°, 464 m], L. Borimecico leg., 27.II.2014, 1 ♂ ( IBSP 211200 ) ; Pereira Barreto, Usina Hidrelétrica de Três Irmãos [S20.66905°, W51.30087°, 310 m], P.A.M. Goldon & S. Guizze leg., X.1990, 1 ♂ ( IBSP 4836 ) ; Pinhalzinho [S22.7827°, W46.59084°, 947 m], P.A.M. Goldon & S. Guizze leg., 4.XI.2008, 1 ♀ ( IBSP 123984 ) ; Pirapora do Bom Jesus [S23.40362°, W46.99886°, 739 m], P.A.M. Goldon & S. Guizze leg., 1–7.III.2013, 1 ♂ ( IBSP 210614 ) ; Primavera, Usina Hidrelétrica Engenheiro Sérgio Motta [S22.4877°, W52.95655°, 252 m], P.A.M. Goldon & S. Guizze leg., 20–25.III.2001, 1 ♀ ( IBSP 38610 ) , same collectors, 23.II.2001, 1 ♂ 2 ♀ ( IBSP 39348 ) ; Usina Hidrelétrica Sérgio Motta, P.A.M. Goldon & S. Guizze leg., I–II.2000, 3 ♂ 7 ♀ ( IBSP 30092 ) , G.R.S. Ruiz leg., same data, 6 ♂ 4 ♀ ( IBSP 30097 ) ; Rio Claro, Floresta Estadual Edmundo Navarro Andrade [S22.41236°, W47.55162°, 623 m], 1.IX.2009, 1 ♂ ( IBSP 146382 ) [S22.40893°, W47.56024°, 623 m], E.O. Machado leg., 16.III.2005, 1 ♀ ( IBSP 51353 ) , F.U. Yamamoto leg., same data, 1 ♂ ( IBSP 51361 ) , R. Yamamoto leg., 6.VI.2005, 1 ♀ ( IBSP 53027 ) , A. Girotti leg., same data, 1 ♂ ( IBSP 53029 ) , 10.XII.2008, 1 ♂ ( IBSP 122264 ) , W.N. Silva leg., same data, 1 ♂ ( IBSP 122265 ) , P.A.M. Goldon & S. Guizze leg., same data, 1 ♂ ( IBSP 122266 ) , D.F. Candiani leg., 10.X.2011, 1 ♀ ( IBSP 163008 ) ; Santana de Parnaíba (S23.44417°, W46.91778°, 748 m), E.G. Achú leg., 15.IV.2003, 1 ♀ ( IBSP 37131 ) ; São Carlos, Campus da UFSCar, Bosque Pinus [S21.98429°, W47.88057°, 856 m], D.F. Candiani leg., XII.2006, 4 ♀ ( IBSP 78890 ) ; São Paulo, Bairro Butantã [S23.56386°, W46.71647°, 739 m], IV.1991, 1 ♂ ( IBSP 4230 ) ; Campus USP [S23.56734°, W46.73241°, 739 m], R. D’Avilla leg., 10.V.2005, 1 ♀ ( IBSP 57263 ) , Instituto Butantan [S23.56718°, W46.71876°, 739 m], A.J. Santos leg., 14.II.2005, 1 ♀ ( IBSP 56972 ) ; Mata do Instituto Butantan , 10.II.2017, 1 ♀ ( IBSP 220688 ) ; Parque dos Príncipes [S23.57302°, W46.77051°, 770 m], R. Martins & R. Bertani leg., 5–12.X.2004, 1 ♂ ( IBSP 48534 ) , G.R.S. Ruiz leg., same data, 1 ♂ ( IBSP 48535 ) , L.A. dos Anjos leg., VII.2004, 1 ♀ ( IBSP 48679 ) , G.P. Perroni leg., same data, 1 ♂ ( IBSP 48680 ) , G.P. Perroni leg., same data, 1 ♂ ( IBSP 48692 ) , G. Perroni leg., same data, 1 ♂ ( IBSP 48695 ) , S. Barros leg., 5–12.X.2004, 1 ♂ ( IBSP 48532 ) ; Parque Estadual da Cantareira, Sede [S23.45468°, W46.63569°, 949 m], R.P. Indicatti leg., 4.III.2001, 1 ♀ ( IBSP 85700 ) ; Rodovia Fernão Dias , km 58 [S23.49391°, W46.55904°, 748 m], VIII.1989, 1 ♀ ( IBSP 4930 ) , same locality [S23.65431°, W46.63239°, 792 m], E.G. Achú leg., 14.IX.2006, 1 ♀ ( IBSP 72914 ) , same locality [S23.65114°, W46.69327°, 761 m], 13.X.2010, 1 ♂ ( IBSP 211411 ) , same locality (S23.5475°, W46.63611°, 778 m), same collector, XI.1991, 1 ♂ ( IBSP 5129 ) ; São Paulo, Campus USP, Reserva Florestal da CUASO [S23.56288°, W46.72973°, 739 m], I. Knysak & R. Martins leg., 4.VIII.1999, 1 ♀ ( IBSP 32943 A) [S23.5475°, W46.63611°, 778 m], V.E. Sandoval leg., I.1991, 1 ♂ 1 ♀ ( IBSP 4063 ) ; São Pedro [S22.55368°, W47.91154°, 551 m], R. Martins & G. Puorto leg., 11.XII.2003, 1 ♂ ( IBSP 41989 ) ; Socorro, Pousada da Gruta do Anjo [S22.60361°, W46.50583°, 801 m], I. Knysak & R. Martins leg., 19–22.XII.2010, 1 ♂ 1 ♀ ( IBSP 213276 ) ; Sorocaba, Mata Fechada da Mata da UNIP [S23.46007°, W47.41937°, 580 m], I. Knysak & R. Martins leg., 14.VI.2008, 1 ♂ ( IBSP 99061 ) ; Tietê [S23.10179°, W47.71468°, 498 m], I. Knysak & R. Martins leg., 22.IV.2003, 2 ♀ ( IBSP 51253 ) ; Tupã [S21.93287°, W50.5049°, 503 m], I. Knysak & R. Martins leg., 16– 20.VII.2008, 1 ♀ ( IBSP 118041 ) ; Valinhos, Fazenda Santo Antônio da Cachoeira (S22.93675°, W46.90834°, 771 m), L. A. dos Anjos leg., IV.2002 – III.2003, 6 ♂ 2 ♀ ( IBSP 78540 ) ; Vargem Grande do Sul [S21.8369°, W46.90131°, 721 m], G.P. Perroni leg., 26.XII.2007 – 5.I.2008, 1 ♂ ( IBSP 99253 ) , G.P. Perroni col. leg., 13.V.2005, 1 ♂ 1 ♀ ( IBSP 63185 ) , G. Perroni leg., 2005, 1 ♀ ( IBSP 119859 ) . Paraná: Altônia, Parque Nacional de Ilha Grande, Margem da Lagoa Xambre [S23.85139°, W53.99234°, 284 m], T.G. Pinheiro leg., 20.IX.2010, 1 ♀ ( IBSP 167263 ) ; Parque Nacional Ilha Grande, Margem da Lagoa Xambrê , T.G. Pinheiro leg., 24.I.2009, 1 ♀ ( IBSP 135489 ) , 1 ♂ ( IBSP 135490 ) , same collector, IV.2009, 1 ♀ ( IBSP 145314 ) ; Londrina, Parque Estadual Mata dos Godoys [S23.43975°, W51.24091°, 581 m], T.G. Pinheiro leg., 18.X.1999, 1 ♀ ( IBSP 38277 ) , same collector, 30.IIII.2000, 1 ♀ ( IBSP 38287 ) , same collector, 30.III.2000, 1 ♀ ( IBSP 38293 ) ; Ponta Grossa [S25.10059°, W50.15808°, 899 m], T.G. Pinheiro leg., 2008, 1 ♂ ( IBSP 145398 ) . Santa Catarina: Florianópolis, Itacorubi , UCAD [S27.53146°, W48.51277°, 10 m], H. Ferrarezzi leg., 5–12.I.2004, 1 ♂ ( IBSP 144326 ) ; São Cristóvão do Sul, Monte Alegre [S27.30277°, W50.42401°, 1060 m], D.F. Júnior leg., 2001–2002, 1 ♂ ( IBSP 141455 ) , G.Q. Romero leg., same data, 1 ♂ ( IBSP 141456 ) ; São Miguel d’Oeste [S26.73727°, W53.51845°, 644 m], J. Casoy leg., III–VIII.2004, 1 ♂ ( IBSP 79203 ) . Rio Grande do Sul: Novo Hamburgo, Lomba Grande [S29.76378°, W51.04019°, 74 m], L.S.P. Trigueiro, C.A.D. Teixeira, L.A.P. Sallet & K.S. Serra leg., XI–XII.2007, 1 ♂ 1 ♀ ( IBSP 98698 ) ; Porto Alegre, Lomba do Pinheiro [S30.10046°, W51.12604°, 96 m], L.S.P. Trigueiro et al., leg., II–III.2005, 2 ♂ ( IBSP 51367 ) ; Santa Maria, Três Barras [S29.65374°, W53.67684°, 122 m], L.S.P. Trigueiro et al., leg., 30.IX.1992, 1 ♂ ( IBSP 122635 ) , same locality [S29.6935°, W53.79118°, 134 m], L.S.P. Trigueiro et al., leg., 27.X.2008, 1 ♂ ( IBSP 214628 ) ; São Borja (S29.07611°, W56.32722°, 74 m), L.S.P. Trigueiro et al., leg., X–XI.2006. IX.2007, 1 ♂ ( IBSP 88679 ) ; São Pedro do Sul [S29.61879°, W54.17773°, 178 m], L.S.P. Trigueiro et al., leg., IX–XI.1988, 1 ♂ ( IBSP 214629 ) ; São Sepé [S30.16818°, W53.56404°, 94 m], L.S.P. Trigueiro et al., leg., 10.II.2000, 1 ♂ 1 ♀ ( IBSP 214630 ) . URUGUAY: Lavallejo: Cerro los Cuervos [S34.27004°, W55.24795], T. Souza et al., leg., 6.XII.2005, 1 ♂ ( IBSP 59513 ) . Montevideo: km 12, Ruta 1 [S34.34822°, W57.15891°], J.M. Bortolin leg., 2001, 2 ♂ ( IBSP 44079 ) , same locality, J. Lopes leg., same data, 2 ♂ ( IBSP 44078 ) , 3 ♀ ( IBSP 44080 ) .

Distribution. Argentina, Brazil, Bolivia, Paraguay, Peru and Uruguay; the records from Cuba and from the southern and western United States ( Valle et al. 2013) are better explained by synanthropic introduction ( Fig. 25 ).

Figures

FIGURE 1. Scanning electron microscopy, Falconina gracilis (Keyserling, 1891), male (IBSP 169327): A cephalothorax and eyes, frontal view; B sternum, ventral view; C chelicerae; D cheliceral teeth. Scale bar A–C. 500 μm; B, 1 mm; D, 300 μm.

FIGURE 2. Scanning electron microscopy, Falconina gracilis (Keyserling, 1891), A–E male (IBSP 169327), F female (IBSP 72914): A left claw, leg I; B left claws, leg IV; C–D tarsal organ; E trichobothria; F left palpal claw. Scale bar C, 20 μm; D–E, 10 μm; A, B, F, 50 μm. Abbreviation TO, tarsal organ.

FIGURE 3. Living specimens of Falconina gracilis (Keyserling, 1891) from Goiânia, Goiás, Brazil: A male (MPEG 37222); B female (MPEG 37222). Photos by A. Anker.

FIGURE 4. Scanning electron microscopy, Falconina gracilis (Keyserling, 1891) male (IBSP 169327), spinnerets:A spinnerets, ventral; B left ALS; C right PLS; D left PMS; Abbreviations Ac, aciniform gland spigot; ALS, anterior lateral spinnerets; AT, anal tubercle; Co, colulus; MaAmp, major ampullate gland spigot; MiAmp, minor ampullate gland spigot; Nu, nubbin; PLS, posterior lateral spinnerets; PMS, posterior median spinnerets; Pi, piriform gland spigot; Tp, tartipore; TrSP, tracheal spiracle. Scale bar A, 200 μm; B–D, 40 μm; C, 50 μm, D, 40 μm.

FIGURE 5. Scanning electron microscopy, Falconina gracilis (Keyserling, 1891), left male palp (IBSP 169327):A ventral view; B retrolateral view; C apex of palp, embolus and conductor details, ventral view. Abbreviations AS, apical spur; C, conductor; CRP, cymbial retrolateral basal process; dVL, dorsal lobe of the ventral lobe of the retrolateral tibial apophysis; E, embolus; EP, embolar process; DL, dorsal lobe of the retrolateral tibial apophysis; ML, median lobe of the retrolateral tibial apophysis; T, Tegulum; TP, tegular process; VL, ventral lobe of the retrolateral tibial apophysis. Scale bar A–B. 500 μm; C. 400 μm.

FIGURE 6. Scanning electron microscopy, Falconina gracilis (Keyserling, 1891), female genitalia, digested (IBSP 72914): A ventral view; B posterior margin of the epigynal plate and copulatory opening; C dorsal view. D same, primary and secondary spermathecae details. Abbreviations CO, Copulatory opening; FD, fertilization ducts; MnEP, Median notch of the posterior margin of epigynal plate; PVP, posterior vulvar plate; S1, primary spermathecae; S2, secondary spermathecae. Scale bar A–C, 300 μm; B–D 100 μm.

FIGURE 7. A–C. Living female of Falconina gracilis (Keyserling, 1891) (MPEG 37220) from Parque Nacional das Emas, Mineiros, Goiás, Brazil, predating on Atta sp. (Formicidae). Photos by A. Anker.

FIGURE 8. Scanning electron microscopy, Falconina gracilis (Keyserling, 1891), left male palp variation: A–C retrolateral view; D–F RTA, retrolateral view; G–I palpal apex, retrolateral view. A, D, G specimen from João Câmara, Rio Grande do Norte, Brazil (MPEG 19973); B, E, H specimen from Goiânia, Goiás, Brazil (MPEG 37222); C, F, I specimen from Belém, Pará, Brazil (MPEG 37228). Abbreviations AS, apical spur; C, conductor; CRP, cymbial retrolateral basal process; E, embolus; EP, embolar process; DL, dorsal lobe of the retrolateral tibial apophysis; ML, median lobe of the retrolateral tibial apophysis; TP, tegular process; VL, ventral lobe of the retrolateral tibial apophysis. Scale bar A, C–E, 200 μm; B, 500 μm; F–I, 100 μm.

FIGURE 9. Falconina gracilis (Keyserling, 1891), A, C, E, female (IBSP 72914), B, D, F, male (IBSP 169327), G, H Falconina gracilis female holotype (NMH1890.7.1 1243): A–B dorsal habitus; C–D carapace anterior view; E female genitalia ventral view; F male palp, ventral and retrolateral view; G, female genitalia ventral view; G original label.

FIGURE 25. Known geographical distribution of Falconina gracilis (Keyserling, 1891). White star represents the type locality of the senior synonym. Blue stars represent the type localities of the junior synonyms. Blue circles represent new records. Green circles represent previous records.