Pseudotorinia marthae ( Boettger, 1902 )

Harzhauser, Mathias & Landau, Bernard, 2023, The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition, Zootaxa 5370 (1), pp. 1-74 : 40-42

Pseudotorinia marthae ( Boettger, 1902)

Figs 4O , 20A–D

* Solarium (Torinia) marthae n. sp. — Boettger 1902: 115.

Solarium (Torinia) marthae n. sp. —Boettger 1907: 136.

Solarium (Nipteraxis) Marthae Boettger — Cossmann 1916: 168, pl. 12, figs 13–15.

Solarium (Nipteraxis) marthae (Boettger) —Zilch 1934: 217, pl. 13, figs 16a–c.

Architectonica (Nipteraxis) marthae Boettger — Korobkov 1955: plate captions, pl. 13, figs 20–22.

Architectonica (Nipteraxis) marthae (Boettg.) — Báldi 1966: 60.

Architectonica (Nipteraxis) cf. marthae Boettg. —Rado & Mutiu 1970: 147, pl. 6, fig. 22.

Architectonica (Calodisculus) planulata (Grateloup) — Bałuk 1975: 119, pl. 13, fig. 6.

Architectonica (Nipteraxis) marthae (Boettger, 1901) — Atanacković 1985: 97, pl. 23, figs 7–10.

martae (Solarium) ‘Boettger’— Bieler & Petit 2005: 49.

marthae (Solarium) Boettger, 1902 — Bieler & Petit 2005: 49.

Architectonica (Solariaxis) kostejana (Boettger, 1907) — Bałuk 2006: 195: pl. 7, figs 6–7 [non Solatisonax kostejana (Boettger, 1907) ].

Architectonica (Nipteraxis) marthae (Boettger, 1901) — Bałuk 2006: 195: pl. 7, fig. 8.

non Architectonica (Nipteraxis) marthae Boettger — Csepreghy-Meznerics 1956: 386, pl. 1, figs 19–21 [= Pseudotorinia grasemanni nov. sp.].

Architectonica (Nipteraxis) martae [sic] (Boettger)—Kojumdgieva in Kojumdgieva & Strachimirov 1960: 93, pl. 29, figs 17a–d [= Pseudotorinia grasemanni nov. sp.].

non Architectonica (Nipteraxia [sic]) marthae Boettger, 1901 —Strausz 1966: 118, pl. 51, figs 5–7 [= Pseudotorinia grasemanni nov. sp.].

non Solarium marthae Boettger — Csepreghy-Meznerics 1969b: 20, pl. 2, figs 19, 23, 26 [= Pseudotorinia grasemanni nov. sp.].

Type material. Lectotype (designated by Zilch 1934: 218): SMF 372695 (XII 2371a), SL: 1.5 mm, MD: 3.3 mm, Senckenberg Museum, Frankfurt/Main, Germany, Coşteiu de Sus ( Romania), illustrated in Zilch (1934: pl. 7, fig. 16), Figs 20C 1 –C 3 .

Illustrated material. NHMW 2023/0267/0011, SL: 2.5 mm, MD: 5.4 mm, Steinebrunn ( Austria), Figs 20A 1 –A 3 . NHMW 2023/0267/0012, SL: 2.4 mm, MD: 5.7 mm, Steinebrunn ( Austria), Figs 20B 1 –B 3 . NHMW 2023/0267/0013, SL: 2.7 mm, MD: 5.7 mm, Steinebrunn ( Austria), Figs 20D 1 –D 3 . NHMW 2012/0229/0049, SL: 5.1 mm, MD: 12.3 mm, Traiskirchen ( Austria), Fig. 4O .

Additional material. 1spec. NHMW2023 /0267/0014,Steinebrunn( Austria) ; 11spec., NHMW 1870 /0049/0044, Coşteiu de Sus ( Romania) .

Description. Small, depressed lentiform shell with almost flat spire. Protoconch hardly emerging, very small of <1 visible whorl (diameter 0.6 mm). Teleoconch of ~3.5 whorls. First teleoconch whorl flat with slightly raised, weakly beaded SSC and UPC with two delicate, beaded MCs. Later whorls with very prominent SSC of tubercular beads, three weaker MCs and prominent UPC. Beaded LPC appearing during second teleoconch whorl. Subsequent whorl attached below LPC. Beads rounded, subquadratic. LPC forming blunt keel; below steeply inclined, faintly concave passing via weak angulation into weakly convex base. Periphery coarsely beaded in basal view. IPC indistinct followed by three weaker, beaded spiral cords, three broader, beaded spiral cords and only slightly stronger PUC. UC prominent, delimited by narrow groove, overhanging umbilicus. Umbilicus moderately wide (~20–26 of MD). Umbilical wall bearing fine axial growth lines. Aperture small, circular with weak CG.

Discussion. The lectotype is a small subadult specimen that, at its early growth stage, has a rather narrow umbilicus, which becomes only slightly wider during ontogeny. The narrow umbilicus separated Pseudotorinia marthae form the otherwise similar species discussed above under Pseudotorinia grasemanni . Pseudotorinia subplicata sensu Landau et al. (2020b) , from the Tortonian of France, differs in its reticulate sculpture ( Landau et al. 2020b: pl. 1, figs 1–2). Pseudotorinia theresae (Semper, 1861) , from the Pliocene of the Mediterraean Sea, has a broader PUC and a prominent ISC ( Chirli 2013; pl. 6, figs 10–15). The Pliocene Mediterranean Pseudotorinia obtusa ( Bronn, 1831) has a higher spire, a much weaker subsutural spiral cord and a more uniform sculpture ( Ferrero Mortara et al. 1984 pl. 15, figs 4a–c). Some Miocene ‘varieties’ of P. obtusa , described by Sacco (1892), such as P. taurotransiens Sacco, 1892 , from the Langhian of Sciolze, might be close to or identical with P. marthae , but the poor illustrations in Sacco (1892: pl. 2, fig. 47) do not allow a clear identification.

Distribution. Known so far only from the middle Badenian (late Langhian) of the Central Paratethys Sea.

Central Paratethys Sea. Middle Miocene (Badenian): Korytnica Basin: Korytnica ( Poland) ( Bałuk 2006); Hrvaćani ( Bosnia and Herzegovina) ( Atanacković 1985); Făget Basin: Coşteiu de Sus ( Romania) ( Boettger 1902).

Figures

FIGURE 1. Descriptive terms used in descriptions of Architectonicidae (modified from Bieler 1993).

FIGURE 3. Descriptive terms used in descriptions of Mathildidae.

FIGURE 4 Protoconchs of Paratethyan Architectonicidae. A. Ammotectonica gregorovae nov. sp., paratype, NHMW 1862/0029/0078c. B. Simplexollata anticollata nov. sp., paratype, NHMW 1854/0035/0331, Lăpugiu de Sus (Romania). C. Simplexollata carocollata (Lamarck, 1822), NHMW 1850/0018/0115, Léognan (France), Burdigalian. D. Simplexollata simplex (Bronn, 1831), NHMW 2013/0078/0109, Baden (Austria).E. Nipteraxis deformatusnov. sp., paratype, NHMW 2010/0004/0456a, Bad Vöslau (Austria). F. Nipteraxis exmoniliferus (Sacco, 1892), NHMW 1907/0323/1959, M̂llersdorf (Austria). G. Solatectonica kostejana (Boettger, 1907), NHMW 2012/0229/0049b, Traiskirchen (Austria). H. Nipteraxis monilifer (Bronn, 1831), NHMW 1846/0037/0427, M̂llersdorf (Austria). I. Solatisonax tavianii nov. sp., NHMW2016/0266/0002, SL: 8.0 mm, MD: 15.0 mm, Lăpugiu des Sus (Romania). J. Solatisonax transversa nov. sp., holotype, NHMW 2016/0177/0341, Lăpugiu de Sus (Roma- nia). K. Granosolarium semilaevis (Sacco, 1892), NHMW 1846/0037/0428d, M̂llersdorf (Austria). L. Heliacus globosus nov. sp., holotype NHMW 1907/0328/1959a, M̂llersdorf (Austria). M. Heliacus miser (Dujardin, 1837), NHMW 2023/0267/0001, Steinebrunn (Austria). N. Pseudotorinia grasemanni nov. sp., paratype, NHMW 2023/0267/0010, Steinebrunn (Austria). O. Pseudotorinia marthae (Boettger, 1902), NHMW 2012/0229/0049, Traiskirchen (Austria). All figures same scale.

FIGURE 20 A1–A3. Pseudotorinia marthae (Boettger, 1902), NHMW 2023/0267/0011, Steinebrunn (Austria). B1–B3. Pseudotorinia marthae (Boettger, 1902), NHMW 2023/0267/0012, Steinebrunn (Austria). C1–C3. Pseudotorinia marthae (Boettger, 1902), lectotype, SMF 372695, Coşteiu de Sus (Romania). D1–D3. Pseudotorinia marthae (Boettger, 1902), NHMW 2023/0267/0013, Steinebrunn (Austria).