Wallabicoris uptoni, Schuh & P. Pedraza, 2010

Schuh, R. T. & P. Pedraza, 2010, Wallabicoris, New Genus (Hemiptera: Miridae: Phylinae: Phylini) From Australia, With The Description Of 37 New Species And An Analysis Of Host Associations, Bulletin of the American Museum of Natural History 2010 (338), pp. 1-118 : 86-87

Wallabicoris uptoni , new species Figure 46; map 4; plate 5

DIAGNOSIS: Recognized by the large size and the elongate slender body form, total length 5.58, ratio length/width 3.73, the pale setae on the dorsum, the quadrate pattern of coloration on the dorsum (pl. 5), and the form of the male genitalia, the secondary endosomal strap with a moderately broad subbasal undulation, a moderately long left paramere, and a lanceolate right paramere (fig. 46). Separated from other large species with an elongate body form as follows: from W. coolabah by the short submedial twist in the secondary endosomal strap and the short left paramere; from W. craspedii by the heavy black setae on the dorsum and the broad subbasal undulation in the secondary endosomal strap in that species; and from W. norsemanius by the longitudinal rather that quadrate color pattern and the short submedial twist in the secondary endosomal strap of that species.

DESCRIPTION: Male: Body greatly elongate, parallel sided, total length 5.52–5.62, l/w 5 3.73. COLORATION (pl. 5): Pronotum unicolorous faded brown, head and pronotum with a medial longitudinal pale stripe; hemelytron mostly pale, endocorium and exocorium with faded brown areas, clavus mostly faded brown; markings on cuneus present as partial infuscation on posterior two-thirds; membrane with elongate, contrasting, fumose marking at extreme base,

(AMNH_PBI 00168791).

remainder weakly fumose over entire area, veins white; venter heavily infuscate, at least on mesopleuron and abdomen; antennal segment 1 dirty yellow, with one or two black medial setae; antennal segment 2 dirty yellow; labium generally infuscate, heavily so apically; hind femora unicolorous faded brown, without black spots; hind tibial spines dark without dark spots at bases. SURFACE AND VESTITURE: Dorsum with reclining simple setae matching background coloration, without sericeous or woolly setae. STRUCTURE: Head: Elongate, projecting anteriorly, ratio wh/lh 5 2.67; antennal fossa with ventral margin 1 diameter above ventral margin of eye; interocular space moderately large, ratio iod/wh 5 0.41; eyes leaving gena moderately exposed in lateral view (hg3/ he20); labium just reaching onto abdomen. GENITALIA (fig. 46): Endosoma: Base moderately long, with an open U-shaped bend; distal half of shaft smoothly curving; primary endosomal strap weakly elongate apically, ratio lae/lsg 2.00; apex of primary endosomal strap nearly straight; secondary endosomal strap very slender, of uniform width from endosomal bend to gonopore, reaching to level of secondary gonopore, with a broad submedial undulation; secondary gonopore seen laterally in lateral view of endosoma. Phallotheca: Smoothly curving on dorsal margin; dorsal surface with a conspicuous keel. Left Paramere: Body elongate, somewhat exceeding pygophore margin; in dorsal perspective closed over about threefourths length; body not distinctly tapering toward apex; anterior process arising at posterior margin of shaft; anterior process directed vertically; posterior process appearing tubular, fingerlike; posterior process with strongly projecting shoulder at base; base of posterior process at about level of paramere body or very weakly elevated. Right Paramere: Body short and broad, ratio lrp/wrp 3.25; body more or less parallel sided; posterior margin without distinct protuberance subapically; body elevated at juncture with base; apex with short fingerlike process.

ETYMOLOGY: Named for M.S. Upton, collector of the only known specimens.

HOSTS: Unknown.

DISTRIBUTION (map 4): Western Nullarbor Plain, Western Australia.

HOLOTYPE: AUSTRALIA: Western Australia: Madura , 31.928 ° S 126.978 ° E, 02 Oct 1968, Key, Upton and Balderson, 13 (AMNH_PBI 00168793) ( ANIC).

PARATYPES: AUSTRALIA: Western Australia: Madura, 31.928 ° S 126.978 ° E, 02 Oct 1968, Key, Upton and Balderson, 13 (00168791) ( AMNH), 13 (00168792) ( ANIC).