Philodryadini Cope, 1886

Arredondo, Juan Camilo, Grazziotin, Felipe G., Scrocchi, Gustavo J., Rodrigues, Miguel Trefaut, Bonatto, Sandro Luís & Zaher, Hussam, 2020, Molecular phylogeny of the tribe Philodryadini Cope, 1886 (Dipsadidae: Xenodontinae): Rediscovering the diversity of the South American Racers, Papéis Avulsos de Zoologia (Pap. Avulsos Zool., S. Paulo) 60, pp. 1-14 : 5

Tribe Philodryadini Cope, 1886

Type-genus: Philodryas Wagler, 1930 by original designation and monotypy.

Type species: Coluber olfersii Lichtenstein, 1823 .

Content: Chlorosoma Wagler, 1830 resurrected; Ditaxodon Hoge, 1958 ; Philodryas Wagler, 1830 ; Xenoxybelis Machado, 1993 resurrected.

Diagnosis: Members of the tribe Philodryadini can be distinguished from the other xenodontine genera by the following combination of characters: hemipenis bilobed, semicalyculate and semicapitate; large body calyces cov- ering both medial and distal portions of the asulcate side of the hemipenial body and lobes ( Figs. 3-4 ); maxillary dentition diacranterian and opisthoglyphous, with a deep groove that opens laterally.

Geographical distribution: most of the Cis-Andean portion of South America, from southern Colombia, Venezuela, Guianas, Ecuador, Peru, Brazil, Bolivia, Paraguay, Uruguay, and Argentina, and in the Trans- Andean portion of the southern Andes in Chile.

Figures

Figure 3. Hemipenes of Chlorosomaviridissimum (MUSM 2403) in Aand B,and Incaspis amaru (FHGO 4749) inC and D. Photographs in Aand C arein sulcate views, and photographs in B and D are in asulcate views.Scale bars:5 mm.

Figure 4. Hemipenis of Philodryas chamissonis (MNHN 3807) in A and B, Tropidodryas serra (MNRJ 7354) in C and D,and Xenoxybelis argenteus (BMNH 1994.7000) in E and F.Photographs in A,C,and E are in sulcate views,and photographs in B, D,and F are inasulcate views.Scale bars:5 mm.