Furochrysa alisae, Makarkin & Perkovsky, 2024

Makarkin, Vladimir N. & Perkovsky, Evgeny E., 2024, Nothochrysinae (Neuroptera: Chrysopidae) from the early Eocene Fur Formation, Denmark, with description of a new genus, Zootaxa 5433 (4), pp. 529-545 : 534-535

Furochrysa alisae sp. nov.

( Figs 4–5 )

Etymology. From the first name of Alisa A. Perkovsky, who found the specimen in collections of the NHMD (an unnumbered part of the former Rettig collection).

Type material. Holotype: MGUH 34321, collected in 2002 by Erwin Rettig and deposited in the NHMD: slightly crumpled and damaged overlapping fore- and hind wings lacking their basal parts.

Type locality and horizon. Denmark: northern Jutland (Region Nordjylland): Mors Island ( Morsø Kommune ): Svalklit. Earliest Eocene, Fur Formation .

Description. Forewing 13.1 mm long (as preserved; estimated complete length ca. 14 mm), ca. 4.5 mm wide. Costal space relatively broad proximally, narrow distally. All preserved subcostal veinlets simple, rather widely spaced proximally, very dense distally. Sc poorly discernible distally, probably rather long (all veinlets figured in Fig. 4A highly likely belong to Sc). RA long, ending on margin markedly before wing apex; its veinlets poorly discernible. Subcostal space narrow; no subcostal crossveins detected. RA space broad, with 16 crossveins: basal crossveins with apices slightly inclined to wing base; most distal crossveins with apices strongly obliquely to wing apex. Anterior trace of RP slightly zigzagged, apically not forked; with 18 pectinate branches: RP1 presumably forked at Psc, both branches again forked once; RP2, RP3 fused proximally (far proximad Psm); RP5 forked twice distally; RP2–RP4, RP6–RP12, RP18 deeply forked once distally; RP13 shallowly forked once distally; RP14– RP17 simple. Basal crossvein between RP, M (1r-m) long, connecting anterior trace of RP, MA near proximal end of im length. M dividing to MA, MP markedly distad origin of RP. MA probably forked distad Psc; MP probably deeply forked at Psc. im broad (length/width ratio 2.4:1), pentagonal. Crossvein between MA, MP (1im) long. Psm poorly developed, zigzagged. Crossvein 2m-cu connected im, CuA located nearly in middle of im (slightly in distal portion). CuA probably with three branches. CuA continuing into well developed, only slightly zigzagged Psc, which continues into outer gradate series of crossveins. Two gradate series of crossveins clearly convergent; inner series with 17 crossveins distad MA; outer series with 14 preserved crossveins distad RP2 (16 when complete). Long crossvein present between these series (labelled ma-rp 1 in Fig. 2A ). CuP incompletely preserved; its anterior branch probably simple; posterior branch probably very fragmentarily preserved. 2icu long. Presumed A1, A2 fragmentarily preserved, simple.

Hind wing ca. 11.5 mm long, ca. 4.0 mm wide. Costal space narrow; preserved subcostal veinlets simple, widely spaced. Sc indiscernible distally. Subcostal space narrow; crossveins not detected. RA long, ending on margin well before wing apex. RA veinlets poorly discernible. RA space moderately broad; crossveins strongly oblique in distal half. RP with 17 branches; RP1, RP3–RP13 forked once; RP2 forked twice, RP14–RP17 simple. MA, MP basally not preserved (it is unknown if MA fuses with PR). MA presumably forked distad Psc. MP presumably fused with CuA for very short distance; forked distally at Psc with both branches simple. Intramedial space broad; proximal crossvein between MA, MP (1im) very long. Psm relatively well developed, zigzagged. CuA presumably pectinate, with three branches. Psc relatively well developed, zigzagged. CuP simple. Crossvein between CuA, CuP (1icu) very long. A1, A2, A3 simple. Two gradate series of crossveins arranged in smooth line: inner series with 15 crossveins distad MA; outer series with 14 crossveins distad MA.

Figures

FIGURE 2. Stephenbrooksia multifurcata Willmann, 1993, wing venation of the holotype based on the part and counterpart.A, forewing; B, hind wing. Scale bar = 2 mm (both to same scale).

FIGURE 4. Furochrysa alisae gen. et sp. nov., holotype MGUH 34321. A, specimen as preserved; B, basal part of wings; C, apical part of wings. Scale bars = 2 mm (A), 1 mm (B, C).

FIGURE 5. The venation of Furochrysa alisae gen. et sp. nov., holotype MGUH 34321. A, forewing; B, hind wing. Scale bar = 2 mm (both to same scale).