Joania cordata ( Risso, 1826 )

Bitner, Maria Aleksandra, Lozouet, Pierre & Cahuzac, Bruno, 2013, Upper Oligocene (Chattian) brachiopod fauna from the Aquitaine Basin, southwestern France and its paleoenvironmental implications, Geodiversitas 35 (3), pp. 579-606 : 590-592

Joania cordata ( Risso, 1826)

( Fig. 6 A-I; Table 6)

Terebratula cordata Risso, 1826: 389 .

Argyrotheca cordata – Logan 1979: 50-54, text-figs 13, 14, pl. 5, figs 1-12. — Taddei Ruggiero 1985: 371, 372, pl. 7, figs 10-14; 1994: 208, pl. 2, fig. 1-3; 1996: 202. — Bitner 1990: 140-143, text-figs 7, 8, pl. 5, figs 1-14, pl. 7, fig. 1; 1993: 150, pl. 4, figs 1-5; pl. 5, figs 1-8. — Bitner & Pisera 2000: 9-10, pl. 2, figs 1-9. — Bitner & Dulai 2004: pl. 3, figs 9, 10. — Bitner & Kaim 2004: 196, 197, figs 2D-K, 3A-E. — Álvarez & Emig 2005: 155- 158, figs 25F, G, 47, 48. — Dulai 2007: 4-5, fig. 4.1-9. Joania cordata – Álvarez, Brunton & Long 2008: 400- 402, fig. 4C-D, 5A-F, 11-13.

MATERIAL EXAMINED. — Peyrère: three complete specimens, three ventral and one dorsal valves. Abesse: seven complete specimens, four ventral and two dorsal valves, MNHN.F.A46526-A46528 and A46551. Estoti: one complete specimen MNHN.F. A46523 and five dorsal valves MNHN.F. A46524 - A46525 . Bélus: four complete specimens, three ventral and four dorsal valves, MNHN.F. A46529 - A46531 .

OCCURRENCE. — fflis species is common in the Neogene of the Central Paratethys and the Mediterranean province, living today in the north-eastern North Atlantic and the Mediterranean Sea ( Logan 1979, 1983, 1988) at the depth of 3 to 600 m ( Logan 2007).

REMARKS ffle genus Joania was recently established by Álvarez et al. (2008) for those Argyrotheca species that have a narrow hinge line, prominent cardinal process and tubercles on the inner margin. ffle specimens from the Aquitaine Basin agree well with those described from the Miocene of the Central Paratethys ( Bitner 1990, 1993; Bitner & Pisera 2000; Bitner &Kaim 2004), differing from the Recent ones described and illustrated by Logan (1979) in having a lower median septum. ffle shell is small, smooth or covered with poorly defined, shallow ribs. ffle large foramen is triangular with a wide pedicle collar supported by a low median septum. ffle inner socket ridges are narrow but high, and the cardinal process is prominent. ffle specimens bear tubercles on the inner margin of both valves.

Figures

FIG. 6. — Joania cordata (Risso, 1826), Upper Oligocene: A-C, dorsal views of complete specimens; A, Estoti, MNHN.F.A46523, B, C, Abesse, MNHN.F.A46526-A46527; D, inner view of ventral valve, Bélus, MNHN.F.A46529; E, F, inner and oblique lateral views of dorsal valve, young individual,Estoti, MNHN.F.A46524; G-I, inner, oblique lateral views of dorsal valve, and enlargement of anterior part to show septum and connection with a loop, Bélus, MNHN.F.A46530. All SEM. Scale bars: A, E, F, H, I, 500 µm; B-D, G, 1 mm.