Oodes (Oodes) tokyoensis Habu, 1956

Guéorguiev, Borislav & Liang, Hongbin, 2020, Revision of the Palaearctic and Oriental representatives of Lachnocrepis LeConte and Oodes Bonelli (Coleoptera: Carabidae), with special account on Chinese species, Zootaxa 4850 (1), pp. 1-89 : 54

16. Oodes (Oodes) tokyoensis Habu, 1956

Oodes helopioides tokyoensis Habu, 1956: 82 (type locality: ‘Shakujii, Nerima-ku, Tokyo Metropolis’).

References.

Oodes (Oodes) helopioides tokyoensis: Habu 1956: 79–80 (identification keys), 82–85 (description; comparison with Oodes helopioides helopioides and Lachnocrepis prolixa ); Habu 1958: 194 (distribution in Japan).

Oodes helopioides tokyoensis: Lafer 1973: 847–849 (taxonomic notes and comparisons with O. helopioides s.str. and O. integer ); Lorenz 1998: 305; Bousquet 2003: 445; Lorenz 2005: 325; Bousquet 2017: 636.

Oodes tokyoensis: Lafer 1989: 204 (diagnostic features); Bousquet 1996: 472 (relationships with O. amaroides ).

Type material. Holotype preserved in the Insect Museum of the National Institute for Agro–Environmental Science, Tsukuba (https://www.naro.affrc.go.jp/org/niaes/type/dbcarabidae/o_tokyoensis.html). Not examined.

Other material examined. None.

TME: none.

Diagnosis. See “Diagnosis” and “Notes” under O. integer .

Description. See Habu (1956: 82–84).

Distribution. Japan (Hokkaido, Honshu).

Notes. We follow the view of Lafer (1973: 849) that O. tokyoensis is best treated as a distinct species, not a subspecies of O. helopioides . The Russian author based his statement on the fact that the easternmost known locality for the latter species comes from the area of Minusinsk Hollow, south–central Siberia. Hence, a vast area between the Minussinsk Hollow and Japan is not populated by either species, and no hybridization zone exists between them.