Hepatidae, Stimpson, 1871

GUINOT, DANIÈLE, TAVARES, MARCOS & CASTRO, PETER, 2013, Significance of the sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura), with new nomina for higher-ranked podotreme taxa, Zootaxa 3665 (1), pp. 1-414 : 286-289

Hepatidae

Hepatus pudibundus (Herbst, 1785) , male 45.0 × 63.0 mm, Martinique (MNHN-B24668)

Osachila lata Faxon, 1893 , male 31.0 × 41.0 mm, Mexico (MNHN-B20782)

Hexapodidae

Hexaplax megalops Doflein, 1904 , numerous males and females, New Caledonia (MNHN-B19244 and B19245); male 12.0 × 17.0 mm, New Caledonia ( MNHN-B19244 )

? Hexapus sexpes ( Fabricius, 1798) , female 10.6 × 18.0 mm (found already dissected), Persian Gulf (MNHN- B 10209)

Paeduma cylindraceum ( Bell, 1859) , male 15.3 × 23.5 mm, holotype (dry, in good condition), with Bell's handwritten labels: “ Amorphopus ” and “ Am: cylindraceus male sign. Gallapagos [sic] Mr Cuming”, Galápagos Is. (OUMNH Zoo. Coll.); female 9.4 × 14.4 mm, west coast of Mexico, Gulf of Tehuantepec ( USNM 170897 )

Parahexapus africanus (Balss, 1922) , male 4.0 × 6.0 mm, Congo (R.O.C.) (MNHN-B10201)

Pseudohexapus platydactylus Monod, 1956 , ovig. female 6.5 × 8.5 mm, Mauritania (MNHN-B9528)

Thaumastoplax anomalipes Miers, 1881 , male 4.0 ×.6.5 mm, females 4.5 × 7.0 mm, 5.0 × 7.0 mm, Guinea (MNHN-B10202); female 5.2 × 8.5 mm, Guinea, Loos Is. (MNHN-B10203); female 4.0 × 6.0 mm, Pointe- Noire (MNHN-B27913)

Hexalaughlia orientalis (Rathbun, 1909) , male 5.5 × 8.0 mm, Japan, Tomaga Shima Lt. (USNM 46392)

Hexapodidae sp., female 4.3 × 3.0 mm, Gabon (MNHN-B27908)

Homolidae H. Milne Edwards, 1837 (see Appendix II)

Homola barbata ( Fabricius, 1793) , male 30 × 22.3 mm, Ibero-Moroccan Gulf (MNHN-B13764)

Homola coriolisi Guinot & Richer de Forges, 1995 , male 21.4 × 19.0 mm, holotype, New Caledonia (MNHN- B16691)

Homola orientalis Henderson, 1888 , males 34.8 × 23.0 mm, 31.6 × 25.2 mm, French Polynesia (MNHN-B22385)

Homola ranunculus Guinot & Richer de Forges, 1995 , male 32.4 × 26.8 mm, holotype, New Caledonia (MNHN- B16723); male 42.6 × 35.5 mm, paratype, New Caledonia ( MNHN-B20272 ) ; female 41.4 × 38.1 mm, paratype, New Caledonia ( MNHN-B19869 )

Homola vigil A. Milne-Edwards, 1880, male 21.0 × 18.0 mm, lectotype, Martinique ( MNHN-B6964 )

Homolax megalops Alcock, 1894 , male 27.6 × 24.5 mm, India, Madras (BMNH 1895.1.2.17)

Homolochunia kullar Griffin & Brown, 1976 , male 26.5 (without rostrum) × 23.0 mm, New Caledonia (MNHN- B16662)

Homologenus levii Guinot & Richer de Forges, 1995 , male 13.0 × 8.5 mm, holotype, New Caledonia (MNHN- B16682); males and females, paratypes, Coral Sea, Chesterfield Is. ( MNHN-B19983 )

Homolomannia sibogae Ihle, 1912 , male 39.2 × 27.0 mm, Philippines (MNHN-B13825)

Ihlopsis tirardi Guinot & Richer de Forges, 1995 , male 20.0 × 17.0 mm, holotype, New Caledonia, île des Pins ( MNHN-B20283 )

Lamoha inflata (Guinot & Richer de Forges, 1981) , female 34.0 × 29.0 mm, Samoa, Apolima Strait (MNHN- B7024)

Latreillopsis gracilipes Guinot & Richer de Forges, 1981 , male 11.5 × 8.0 mm, holotype, New Caledonia (MNHN- B7032 )

Moloha majora (Kubo, 1936) , male 51.0 × 38.0 mm, Japan, Sagami Bay (MNHN-B24798); ovig. female 43.8 × 38.6 mm, Sagami Bay (MNHN-B24798)

Paromola bathyalis Guinot & Richer de Forges, 1995 , male 97.0 × 75.6 mm, holotype, New Caledonia (MNHN- B20105); male 72.0 × 48.0 mm, paratype, New Caledonia ( MNHN-B19898 )

Paromola cuvieri ( Risso, 1816) , male 58.0 × 48.0 mm, Gulf of Guinea (MNHN-B13611)

Homolodromiidae

Homolodromia bouvieri Doflein, 1904 , ovig. female 33.0 × 26.0 mm, Madagascar (MNHN-B28940)

Homolodromia kai Guinot, 1993 , ovig. female 25.0 × 18.0 mm, holotype, Indonesia, Kai Is. ( MNHN-B22845 ); female 12.4 × 8.3 mm, same data ( MNHN-B22704 )

Homolodromia paradoxa A. Milne-Edwards, 1880, female 24.0 × 15.0 mm, Bahamas (RMNH)

Dicranodromia doederleini Ortmann, 1892 , ovig. female, Japan, Sagami Bay, lectotype, selected by Guinot (1995) (MZS 567) (see Komai 1999)

Dicranodromia karubar Guinot, 1993 , 18.5 × 12.4 mm, holotype, Indonesia, Tanimbar I. ( MNHN-B22846 )

Dicranodromia felderi Martin, 1990 , male 23.0 × 14.0 mm, Caribbean Sea, in the vicinity of San Andrés I. (USNM)

Dicranodromia foersteri Guinot, 1993 , male 18.0 × 12.0 mm, holotype, Coral Sea, Chesterfield Is. (MNHN- B22700)

Dicranodromia mahieuxii A. Milne-Edwards, 1883, male 9.0 × 6.5 mm, holotype, France, Bay of Biscay (MNHN- B 21681)

Hymenosomatidae

Amarinus lacustris ( Chilton, 1882) , male 4.6 × 5.8 mm, female 4.7 × 4.8 mm, juv. 2.0 × 2.0 mm, New Zealand, Auckland, Oratia stream ( ZRC2010.0238 )

Amarinus laevis (Targioni-Tozzetti, 1877) , female 5.7 × 6.1 mm, Australia, SE Queensland, Hervey Bay, Mary River (QM W9745)

Cancrocaeca xenomorpha Ng, 1991 , male 3.9 × 4.6 mm, paratype, Indonesia, Sulawesi, Maros, Lubang Battu Neraka cave ( MNHN-B24450 ); ovig. female 4.1 × 5.2 mm, Indonesia, Sulawesi, Selatan , Gua Samanggi ( ZRC 2007.0118 )

Elamena truncata ( Stimpson, 1858) , male, Taiwan, Taipei ( ZRC 1999.2062 )

Guaplax denticulata Naruse, Ng & Guinot, 2008 , female 3.4 × 4.1 mm, paratype, Indonesia, Timur, Kalimantan, Gua Laowacling ( MNHN-B30161 )

Halicarcinus mexicanus (H. Milne Edwards, 1853) , male 7.3 × 7.7 mm, holotype (dry), Mexico ( MNHN-B655 )

Halicarcinus planatus (Fabricius, 1775) , male 17.0 × 21.0 mm (dissected), ovig. female 15.0 × 20 mm, Kerguelen Is., Bouvier det. (MNHN-B25953); male 15.0 × 17.3 mm (dissected), several males and females, South Pacific Ocean, Campbell I. (MNHN-B25960)

Hymenicoides robertsi Naruse & Ng, 2007 , males 4.0 × 5.8 mm– 4.9 × 5.7 mm, 2 ovig. females 3.8 × 4.4, 4.2 × 4.8 mm, paratypes, Myanmar, Raknine State , market on Kaladan River ( MNHN-B30393 )

Hymenosoma orbiculare Desmarest, 1823 , male 23.2 × 23.9 mm, lectotype ( MNHN-B10998 ), female 12.5 × 11.1 mm, paralectotype, Cape of Good Hope ( MNHN-B649 ) (both in dry condition)

Limnopilos sumatranus Naruse & Ng, 2007 , male 4.6 × 5.1 mm, female 5.4 × 6.3 mm, paratypes, Indonesia, Sumatra ( MNHN-B30395 )

Neorhynchoplax mangalis (Ng, 1988) , ovig. female 3.6 × 3.3 mm, Singapore (ZRC 1993.6752)

Neorhynchoplax prima Ng & Chuang, 1996 , holotype, male, Indonesia, Pulau Bintan ( ZRC 1994.4233 )

Odiomaris estuarius Davie & Richer de Forges 1996 , male 4.6 × 4.3 mm, holotype, New Caledonia, Dumbea estuary ( MNHN-B25278 ), female 5.4 × 5.0 mm, paratype, same data ( MNHN-B25275 )

Odiomaris pilosus (A. Milne-Edwards, 1873), 1 male 17.3 × 19.8 mm, New Caledonia, Rivière Napoémien, Poindimié ( MNHN-B7908 ) ; 1 male 20.0 × 22,6 mm, New Caledonia ( MNHN-B26146 ) ; several specimens (one male 11.0 × 14.0 mm dissected by M. Tavares in January 2011), New Caledonia, Wé Waalu River , G. Marquet det. ( MNHN-B32604 ) ; 3 ovig. females 9.5 × 10.2 – 10.1 × 11.2 mm, 1 young male, New Caledonia, Rivière de Canala ( MNHN-B32610 )

Sulaplax ensifer Naruse, Ng & Guinot, 2008 , ovig. female 5.0 × 6.5 mm, paratype ( ZRC 2007.0119 ); female 4.0 × 5.0 mm, Indonesia, Sulawesi, Muna I. ( MNHN-B30396 )

Trigonoplax unguiformis (De Haan, 1839) , female 10.4 × 14.5 mm, 2 other females,? Japan ( USNM 13714 ); 1 male 6.1 × 7.3 mm, Western Australia , Passage I., 19 m, coral, sponge and some sea grass, A. B. Williams det. 1981 ( USNM 184973 )

Inachidae (measurements including rostrum)

Achaeus cranchii Leach, 1817 , male16.0 × 9.8 mm, Morocco, Skhirat (MNHN-B30688)

Anisonotus curvirostris A. Milne-Edwards, 1879, male 29.0 × 16.0 mm, holotype (present designation) (dry) ( MNHN-B4403 )

Anomalothir furcillatus ( Stimpson, 1871) , male 24.8 × 18.9 mm, Barbados (dry), “Blake” (MNHN-B3969); female 34.8 × 9.5 mm, Dominica, “Blake” (MNHN-B17810)

Capartiella longipes ( Capart, 1951) , male 8.2 × 6.5 mm, Dahomey (= Benin) (MNHN-B19590); ovig. female 9.4 × 8.1 mm, Nigeria (MNHN-B19602)

Coryrhynchus lobifrons (Rathbun, 1894) , male 20.0 × 11.9 mm, Mexico, Baja California ( MNHN-B20836 )

Coryrhynchus riisei (Stimpson, 1860) , male 13.5 × 10.1 mm (dry), as Podochele Reisei, Guadeloupe (MNHN- B4402)

Cyrtomaia cornuta Guinot & Richer de Forges, 1988 , male 43.6 × 48.0 mm, New Caledonia (MNHN-B20218)

Ericerodes gracilipes ( Stimpson, 1871) , male 9.3 × 6.3 mm, female 8.8 × 6.0 mm, Gulf of Mexico, ”Blake” ( MNHN-B20484 )

Ericerodes hemphillii (Lockington, 1877) , male 15.1 × 10. mm, Mexico, Baja California ( MNHN-B20805 ); male 16 × 11 mm, as Podochela tenuipes Rathbun, 1894 , Mexico, Baja California ( MNHN-B20487 )

Inachus angolensis Capart, 1951 , male 28.0 × 26.6 mm, ovig. female 27.0 × 26.2 mm, Gabon (MNHN-B17412)

Inachus dorsettensis ( Pennant, 1777) , male 20.0 × 18.0 mm, France, Saint Martin de Ré (MNHN-B17337)

Inachus phalangium (Fabricius, 1775) , male 19.5 × 15.1 mm, West Africa (MNHN-B17358)

Metoporhaphis calcarata (Say, 1818) , male cl 19.0 mm (dry), Gulf of Mexico ( MNHN-B4396 )

Podochela grossipes Stimpson, 1860 , male, 13.0 × 10 mm (dry), Saint Lucia (MNHN-B557)

Podochela macrodera Stimpson, 1860 , female 10.0 × 8.0 mm, Guadeloupe (dry) (MNHN-B4401)

Inachoididae (measurements including exposed pleurites 5–8 and rostum)

Anasimus latus Rathbun, 1894 , male 20. 4 × 15.3 mm, female 18.4 × 14.0 mm, French Guiana (MNHN-B17807)

Collodes leptocheles Rathbun, 1894 , male 17.5 × 13.2 mm, Gulf of Mexico ( MNHN-B14055 )

Leurocyclus tuberculosus (H. Milne Edwards & Lucas, 1842) , 45.0 × 50.0 mm, 46.8 × 52.4 mm, syntypes of Salacia tuberculosa (both in dry condition), Chile (MNHN-B3974); juv. 6.0 × 5.5 mm, type of Salacia gracilipes A. Milne-Edwards & Bouvier, 1923 ( Microrhynchus gracilipes A. Milne-Edwards MS), Rio de la Plata (MNHN-B9484); male 33.6 × 36.8 mm, Uruguay,? Leurocyclus gracilipes A. Milne-Edwards & Bouvier, 1902 (MNHN-B9366)

Paradasygyius depressus ( Bell, 1835) , 1 male, Mexico, Sonora ( MNHN-B19502 ) ; 2 specimens, same data ( MNHN-B19503 ) ; 3 specimens, Mexico, Bahía Santa Inés ( MNHN-B20818 )

Paulita tuberculata ( Lemos de Castro, 1949) , male 19.1 x 19.7 mm, French Guiana, Durand coll. (MNHN- B 19506) ; female 17.0 x 17.2 mm, same data ( MNHN-B19511 ) : male, French Guiana, GREEN 0 trawling cruise, Le Loeuff & von Cosel coll. ( MNHN-B28820 ) ; 2 males, 1 ovig. female ( MNHN-B28821 ) ; Brazil, Recife , 1 male, 2 incomplete specimens ( MNHN-B19508 ) ; 1 female ( MNHN-B19507 ) ; 1 male 21.0 x 21.8 mm ( MNHN-B19509 )

Pyromaia tuberculata (Lockington, 1877) , male 17.0 × 12.2 mm, Baja California (MNHN-B9290)

Stenorhynchus debilis (Smith, 1871) , several males and females, Clipperton I. (MNHN)

Stenorhynchus lanceolatus (Brullé, 1837) , male 34.0 × 16.0 mm, female 37.0 × 13.0 mm, several other specimens, West Africa, Guinean Trawling Survey II (MNHN-B21437)

Latreilliidae

Eplumula phalangium (De Haan, 1839) , 2 males, 1 female, Taiwan ( MNHN-B28470 )

Latreillia metanesa Williams, 1982 , 2 males, 1 female, Vanuatu ( MNHN-B28226 )

Latreillia valida De Haan, 1839 , cl 16 mm, Japan (MNHN)

Leucosioidea

Alox antheos C.G.S. Tan & Ng, 1995 , female 8.4 × 6.7 mm, holotype, Indonesia, Aru Is. ( MNHN-B19048 )

Arcania brevifrons Chen, 1989 , male 20.0 × 18.5 mm, holotype, Philippines ( MNHN-B18079 )

Coleusia signata (Paul'son, 1875), male 24.0 × 21.0 mm, female 25.5 × 22.2 mm, Suez Canal (MNHN-B31894)

Ebalia cariosa (Stimpson, 1860) , male 9.2 × 10 mm, Florida, Marco (MNHN-B19851)

Ebalia lacertosa Nobili, 1906 , 9 males, 10 females, syntypes, Red Sea ( MNHN-B19927 )

Euclosia concinna Galil, 2003 , male cl 26.0 mm, holotype, New Caledonia ( MNHN-B28794 )

Euclosia unidentata (De Haan, 1841) , male 29.0 × 27.0 mm, Japan, Tokyo ( MNHN-B16965 )

Ilia nucleus ( Linnaeus, 1758) , male 27.0 × 26.0 mm, Greece, Mytilene, Port Vera (MNHN-B19975)

Iphiculus spongiosus Adams & White, 1849 , male 10.0 × 15.0 mm, Philippines (MNHN-B18103)

Pariphiculus mariannae (Herklots, 1852) , male 26.0 × 23.0 mm, Philippines (MNHN-B18088)

Leucosia anatum (Herbst, 1783) , male 21.0 × 20.0 mm, New Caledonia, (MNHN-B17525)

Myra curtimana Galil, 2001 , male cl 26.0 mm, New Caledonia (MNHN-B21181)

Oreotlos etor C.G.S. Tan & Richer de Forges, 1993 , male 5.5 × 7.5 mm, holotype, Coral Sea, Chesterfield Is. ( MNHN-B24848 )

Oreotlos speciosus Chen, 1989 , female 4.6 × 6.9 mm, holotype, Philippines ( MNHN-B18188 )

Philyra pisum De Haan, 1841 , male 21.0 × 20,5 mm, Japan (MNHN-B16982)

Pseudomyra mbizi Capart, 1951 , male 25.0 × 20.0 mm, Congo (R.O.C.), Pointe-Noire (MNHN-B7570)

Speloeophorus pontifer ( Stimpson, 1871) , syntype of Speloeophorus triangulus A . Milne-Edwards, 1880, damaged, Florida, Charlotte Harbor ( MNHN-B17042 )

Urnalana elata (A. Milne-Edwards, 1873), 3 males, 1 female, syntypes (dry), New Caledonia ( MNHN-B174 )

Urnalana foresti (Chen, 1989) , female 10.0 × 9.1 mm, holotype, Philippines ( MNHN-B18124 )

Urnalana margaritata (A. Milne-Edwards, 1873), 2 males, 2 females, syntypes (dry), New Caledonia (MNHN- B181 )

Litocheiridae

Litocheira bispinosa Kinahan, 1856 , male 8.2 × 10.0 mm, female 8.5 × 11.0 mm, Australia, Tasmania (MNHN- B17754)

Lyreididae

Lyreidus brevifrons Sakai, 1937 , male 49.4 × 23.4 mm, New Caledonia (MNHN-B24319)

Lyreidus tridentatus De Haan, 1841 , male 37.1 × 20.7 mm (MNHN-B12320); female 30.6 × 17.5 mm, Philppines (MNHN-B13373)

Lysirude channeri (Wood-Mason, 1885) , male 25.5 × 17.4 mm, female 33.7 × 20.5 mm, Philippines (MNHN- B 11562)

Macrophthalmidae

Ilyograpsus vanninii Sawada, Hosogi & Sakai, 2005 , male 3.8 × 4.6 mm, Madagascar (MNHN-B6374).

Macrophthalmus (Venitus) latreillei ( Desmarest, 1822) , male 38.0 × 51.0 mm, Madagascar (MNHN-B12614)

Majidae

Maja brachydactyla Balss, 1922 , male 40.0 × 31.0 mm, France, Roscoff (MNHN-B19847) (see Sotele et al. 2008)

Maja squinado (Herbst, 1788) , male 36.0 × 30.0 mm, France, Roscoff (MNHN-B6705)

Prismatopus albanyensis (Ward, 1933) , male 26.0 × 14.0 mm, holotype, Indonesia ( MNHN-B17817 )

Prismatopus harmandi (Bouvier, 1906) , male 46.0 × 24.5 mm, holotype, Japan, Tokyo ( MNHN-B22330 )

Prismatopus longispinus (De Haan, 1839) , female 23.0 × 27.4 mm, Japan (MNHN-B17853)

Mathildellidae

Mathildella maxima Guinot & Richer de Forges, 1981 , male 26.0 × 35.0 mm, holotype, French Polynesia, Tuamotu Archipelago, Mururoa (MNHN-B6812); 2 males 18.1 × 22.7 mm, 23.1 × 29.2 mm, 2 females 16.3 × 20.5 mm, 17.0 × 21.0 mm, Philippines, Balicasag I. ( ZRC 2004.0711 )

Beuroisia duhameli Guinot & Richer de Forges, 1981 , male 58,3 × 83.6 mm, holotype, Saint Paul and Amsterdam Is. ( MNHN-B6809 )

Beuroisia major ( Sakai, 1978) , male 64.0 × 87.0 mm, paratype, North Pacific, Kinmei Seamount (T. Sakai Collection)

Beuroisia manquenei Guinot & Richer de Forges, 1981 , male 36.4 × 50.7 mm, holotype, French Polynesia, Tuamotu Archipelago , Mururoa ( MNHN-B6811 )

Matutidae

Ashtoret lunaris (Forsskål, 1775) , male 40.0 × 59.0 mm, Suez Gulf ( MNHN-B13529 )

Ashtoret sanguiannulata Galil & Clark, 1994 , male cl 48.0 mm, holotype, Madagascar ( MNHN-B21359 )

Menippidae

Menippe nodifrons Stimpson, 1859 , male 39.0 × 56.0 mm, Congo (R.O.C.) (MNHN-B7737); M. mercenaria (Say, 1818) , male, Florida, Key West (MNHN-B9500)

Mictyridae

Mictyris longicarpus Latreille, 1806 , male cl 26.1 mm (found dissected), Australia, Port Jackson (MNHN-B13254)

Mictyris platycheles H. Milne Edwards, 1852 , male 13.3 × 12.3 mm, Australia ( MNHN-B16086 ); male ca 14.0 × 130 mm, Australia, Tasmania ( MNHN-B29599 )

† Necrocarcinidae

† Necrocarcinus labeschii (Deslongchamps, 1835) , Cenomanian, Le Mans, Sables du Perche Formation, one carapace 19.0 × 24.0 mm (MHN LM 3808 ). Albian, Cambridge, five carapaces ( MNHN); Lower Cenomanian, France, Normandy, Seine-Maritime, one carapace (G. Breton Colln); Middle Albian , France, Pas-de-Calais , Wissant , two carapaces (G. Breton Colln)