Anapausis fuscinervis (Edwards)

Amorim, Dalton De Souza & Balbi, Maria Isabel P. A., 2006, A review of Anapausis Enderlein (Diptera: Scatopsidae) in the Neotropical Region, with four new species and comments on the phylogeny of the genus, Zootaxa 1300, pp. 1-29 : 20-22

Anapausis fuscinervis (Edwards)

( Figs. 28–36 )

Material analyzed. HOLOTYPE, S. Chile, Llanquihue Prov., Casa Pangue, 4–10.xii.1926, F. & M. Edwards / B.M. 1927–63. Other specimens: 1 male, Puerto Puyuhuapi, Sud Chile, 14­11­39, G.H. Schwabe; 1 female, Chile, El Naranjo, Tilana, Coquimbo, x.1967, L.E. Peña col. (Natural History Museum, London); 1 male and 1 female, Chiloe, Isla Chiloe, Ahoni Alto, September 1988, Luis E. Peña col. ( CMNH).

The original description of A. fuscinervis and the redescription by Amorim & Haenni (1996) allow correct identification, especially with the illustration of the terminalia of the male holotype. New specimens have been found, allowing a more precise description of the male terminalia and for the first time the illustration of the female terminalia, with some new features added.

Redescription

Male. Similar to A. clivicola sp. nov. except as follows. Setae regularly distributed between the ommatidia ( Fig. 28 ). Maxillary palpus well developed, without sensory pit ( Fig. 29 ). Labella short and slender ( Fig. 30 ). Measurements (in mm): body length, 1.92; wing length, 1.54; wing width, 0.64. Proepisternals, 33–34; proepimerals 12–17; spiracular sclerite setae, 12–17; anepisternals, 16–23; anterior katepisternals, 6–10; mesepimerals 5–6/4–5; merals, 6–7; subspiraculars, 5.

As in A. clivicola , A. fuscinervis presents a cleft on the anteroventral corner of the katepisternum ( Fig. 31 ). The wing ( Fig. 32) has a considerably long, although interrupted, M1. There are setae only on the wing cells M4, CuA2, and anal lobe. The male terminalia are complex ( Fig. 33 ). Sternite 6 has unique modifications on its posterior margin, with a pair of mesal projections covered with scale­like projections and a mesal “pocket”. Sternite 7 is present as a pair of rather large lateral plates, each with an inner projection. There is no evident trace of sternites 8+9, but at most present a slender strip. The gonocoxites seem highly reduced and modified; the gonostyles are digitiform, with a wide base. The aedeagus is tubular distally, not too long, with the anterior pseudopenis projection basally, between the plates of sternite 7. Behind the aedeagus ventrally to the anal membranes, there is a pair of setose sclerites that must be interpreted as the completely divided sternite 10; each sclerite has an inner projection bifid on the apex, and an outer setose, rounded projection, apparently connected mesally behind the apex of the aedeagus. Tergite 8+9 is obvious dorsally, with a pair of well­sclerotized spiracles, although not very conspicuous, and a pair of ventroposterior lobes on tergite 8+9.

Female ( Figs. 34–36 ). Gonapophyses 8 absent, as in A. clivicola sp. nov. and A. mourei sp. nov. Sternite 8 rather short, not projected posteriorly. Tergite 8+9 characteristic, with pair of posterior projections with mesal cleft and dark, short, well­sclerotized mesal tooth. Inner bursae of sternite 9 present.

Figures

FIGURE 31. Anapausis fuscinervis (Edwards). Thorax.

FIGURE 33. Anapausis fuscinervis (Edwards). Male terminalia, ventral view.

FIGURES 28 – 30. Anapausis fuscinervis (Edwards). 28. Detail of ommatidia. 29. Maxillary palpus. 30. Labella.

FIGURES 34 – 36. Anapausis fuscinervis (Edwards). Female terminalia. 34. Ventral view. 35. Dorsal view. 36. Lateral view.