Teruelius intertidalis (Lourenço, 1999) Lowe & Kovařík, 2019

Lowe, Graeme & Kovařík, František, 2019, Review of Grosphus Simon 1880 with description of Teruelius gen n a new buthid genus from Madagascar (Scorpiones Buthidae), Euscorpius 281, pp. 1-128 : 61

Teruelius intertidalis (Lourenço, 1999) comb. n.

1♀ (holotype, Figs. 207 , 222 , 495–515 ), Toliara Province, 3.5 km north of Tulear , IV. 1998, leg. N. Lutzmann ( ZMUH) ; 1♂ 1♀ ( Fig. 85 ), No. 1485 ( FKCP, GLPC) .

Figures

Figures 196–210. Female basal pectinal teeth of representative Grosphus and Teruelius gen. n. G. hirtus (196–198), G. madagascariensis (199), G. madagascariensis, paratype of G. mandena(200), T. ankarafantsika (201),T. ankarana (202), T. annulatus (203),T. bistriatus (204), T. flavopiceus (205), T. grandidieri (206), T. intertidalis (207), T. limbatus (208), T. mahafaliensis (209), T. feti (210).

Figures 211–226. Ventral tarsal setation of legs III or IV in Grosphus and Teruelius gen. n. G. goudoti (211), G. hirtus (212), G. hirtus (G. garciai) (213), G. madagascariensis (214), G. madagascariensis (G. mandena) (215), Teruelius ankarana (216), T. flavopiceus (217), T. annulatus (218), T. ankarafantsika (219–220), T. bistriatus (221), T. intertidalis (222), T. grandidieri (223), T. mahafaliensis (224), T. limbatus (225), T. feti (226).

Figures 495–499. Teruelius intertidalis, female holotype. Figures 495–496. Habitus in dorsal (495) and ventral (496) views, and original labels. Figures 497–499. Metasoma in lateral (497), ventral (498) and dorsal (499) views. Scale bars: 10 mm.

Figures 500–511. Teruelius intertidalis, female holotype. Figures 500–508. Pedipalp chela, dorsal (500), external (501) and ventrointernal (502) views; pedipalp patella, dorsal (503), external (504) and ventral (505) views; pedipalp femur and trochanter, internal (506) and dorsal (507) views; pedipalp chela, movable finger dentate margin (508). Figure 509. Carapace and tergites I–III. Figure 510. Sternopectinal region. Figure 511. Telson lateral view.

Figures 512–515. Teruelius intertidalis, female holotype. Left legs tibia, basitarsus and telotarsus, retrolateral views. Leg I (512), leg II (513), leg III (514), leg IV (515).

Figures 71–85. Hemispermatophores and capsule regions of Teruelius gen n. Multi-panel figures show: whole hemispermatophore; whole hemispermatophore and/or capsule with flagellum; capsule region in convex (or convex compressed), anterior and posterior views (panels in left to right sequence). Right hemispermatophores. Figure 71. T. ankarana, whole hemispermatophore. Scale bar: 4 mm. Figure 72. T. ankarana, capsule, left mirrored, Forêt d’Ankavanana, FMNH. Scale bar: 500 μm. Figure 73. T. grandidieri, whole hemispermatophore (flagellum truncated). Scale bar: 4 mm. Figure 74. T. grandidieri, capsule, Antsakabe River, FMNH. Scale bar: 500 μm. Figure 75. T. flavopiceus, whole hemispermatophore. Scale bar: 2 mm. Figure 76. T. flavopiceus, capsule, Madagascar, GLPC, FKCP. Scale bar: 500 μm. Figure 77. T. annulatus, capsule, Tsimanampetsotsa National Park, GLPC, FKCP. Scale bar: 500 μm. Figure 78. T. ankarafantsika, capsule and flagellum. Scale bar: 400 μm. Figure 79. T. ankarafantsika, capsule, Forêt d’Ankavanana, FMNH. Scale bar: 200 μm. Figure 80. T. ankarafantsika, capsule, Réserve Forestière de l’Ankarafantsika, FMNH. Scale bar: 200 μm. Figure 81. T. olgae, capsule, Itampolo village, FMNH. Scale bar: 500 μm. Figure 82. T. limbatus, whole hemispermatophore (flagellum truncated). Scale bar: 2 mm. Figure 83. T. limbatus, capsule, Forêt d’Ianasana, FMNH. Scale bar: 500 μm. Figure 84. T. mahafaliensis, capsule views, Zombitse-Vohibasia National Park, GLPC, FKCP. Scale bar: 500 μm. Figure 85. T. intertidalis, capsule, Madagascar, GLPC, FKCP. Scale bar: 500 μm.