Pronephrium philippinum (C. Presl) S.E. Fawc. & A.R. Sm.

Fawcett, Susan & Smith, Alan R., 2021, A Generic Classification of the Thelypteridaceae, Fort Worth, Texas, USA: BRIT Press : 72

publication ID	https://doi.org/ 10.17348/jbrit.v15.i2.1206
persistent identifier	https://treatment.plazi.org/id/03B787F6-FFBB-9B18-604A-795BFD16FF6C
treatment provided by	Donat
scientific name	Pronephrium philippinum (C. Presl) S.E. Fawc. & A.R. Sm.
status	comb. nov.

Treatment

Pronephrium philippinum (C. Presl) S.E. Fawc. & A.R. Sm. View in CoL , comb. nov.

— Physematium philippinum C. Presl, Epimel. Bot. 34. 1851.— Nannothelypteris philippina (C.Presl) Holttum — Thelypteris philippina (C. Presl) Ching

Constituent species.—* Pronephrium affine (Blume) C. Presl ; P.amboinense (Willd.) Holttum ; P.amphitrichum Holttum ; P. aquatiloides (Copel.) Holttum ; * P. aoristisorum (Harr.) S.E. Fawc. & A.R. Sm. ; P. borneense (Hook.) Holttum ; P.camarinense (Holttum) S.E. Fawc. & A.R. Sm. ; * P.celebicum (Baker) Holttum ; P. clemensiae (Copel.) Holttum ; P.debile (Baker) Holttum ; P. exsculptum (Baker) Holttum ; P.firmulum (Baker) Holttum ; P. giluwense Holttum ; * P. granulosum (C. Presl) Holttum ; P.hewittii (Copel.) Holttum ; * P.hosei (Baker) Holttum ; P. inaequilobatum (Holttum) S.E. Fawc. & A.R. Sm. ; P. kjellbergii Holttum ; P. lineatum (Blume) C. Presl ; * P. menisciicarpon (Blume) Holttum ; P. merrillii (Christ) Holttum ; P. millarae Holttum ; P. minahassae Holttum ; P. moniliforme (Tagawa & K. Iwats.) Holttum ; P. murkelense (M. Kato) S.E. Fawc. & A.R. Sm. ; P. nervosum (Fée) S.E. Fawc. & A.R. Sm. ; P. palauense (Hosok.) Holttum ; * P. peltatum (Alderw.) Holttum ( Fig. 7C View FIG ); P. peramalense Holttum ; P. philippinum (C. Presl) S.E. Fawc. & A.R. Sm. ; * P. rhombeum (Christ) Holttum ; P.samarense (Copel.) Holttum ; P.simillimum (C. Chr.) Holttum ; P.solsonicum Holttum ; P.thysanoides Holttum ; P.trachyphyllum Holttum ; * P.xiphioides (Christ) Holttum

Excluded species.—Several species included in Pronephrium sect. Dimorphopteris by Holttum (1982) are excluded here. Pronephrium articulatum , distributed in Sri Lanka, India, and southern China, is morphologically distinctive and may be nested within Pseudocyclosorus ( Patel et al. 2019a) . Several species are transferred to Sphaerostephanos ; all other members of Pronephrium sensu Holttum not listed as constituents of Pronephrium s.s. (our new sense) here are treated in Abacopteris , Grypothrix , or Menisciopsis .

Species transferred to Sphaerostephanos .— Pronephrium bakeri (Harr.) Holttum ; * P. beccarianum (Ces.) Holttum ; * P. brauseanum Holttum ; P. bulusanicum (Holttum) Holttum ; P. diminutum (Copel.) Holttum ; * P. glandulosum (Blume) Holttum ; P. melanophlebium (Copel.) Holttum ; P. micropinnatum Holttum ; * P. pentaphyllum (Rosenst.) Holttum ; * P. scopulorum Holttum ; * P. womersleyi Holttum

Incertae sedis.— Pronephrium acanthocarpum (Copel.) Holttum was recognized by Holttum in subg. Menisciopsis . It has setose sporangia, which do not occur in members of that genus as treated here, but are common in species of Abacopteris . Its short stature, trifoliolate laminae, and proliferous buds, however, agree with Grypothrix . Careful study of the type is needed to determine its taxonomic placement.