Sphaerostephanos womersleyi (Holttum) S.E. Fawc.& A.R. Sm.

Fawcett, Susan & Smith, Alan R., 2021, A Generic Classification of the Thelypteridaceae, Fort Worth, Texas, USA: BRIT Press : 82-83

publication ID	https://doi.org/ 10.17348/jbrit.v15.i2.1206
persistent identifier	https://treatment.plazi.org/id/03B787F6-FFB5-9B16-604A-7EC4FDE9FABC
treatment provided by	Donat
scientific name	Sphaerostephanos womersleyi (Holttum) S.E. Fawc.& A.R. Sm.
status	comb. nov.

Treatment

Sphaerostephanos womersleyi (Holttum) S.E. Fawc.& A.R. Sm. View in CoL , comb. nov.

— Pronephrium womersleyi Holttum,Blumea View in CoL 20:108. 1972.

Constituent species.— Sphaerostephanos acrostichoides (Desv.) Holttum View in CoL ; S. adenostegius (Copel.) Holttum View in CoL ; S. alatellus (Christ) Holttum View in CoL ; S. albosetosus (Copel.) Holttum View in CoL ; * S. alcasidii Holttum View in CoL ; S. alpinus Holttum View in CoL ; S. alticola Holttum View in CoL ; S. angustibasis Holttum View in CoL ; S. angustifolius (C. Presl) Holttum View in CoL ; * S. appendiculatus (Blume) Holttum View in CoL ; S. aquatilis (Copel.) Holttum View in CoL ;* S. arbuscula (Willd.) Holttum View in CoL ; * S. archboldii (C. Chr.) Holttum View in CoL ; S. arfakianus (Baker) Holttum View in CoL ; * S. atasripii (Rosenst.) Holttum View in CoL ; S. austerus (Brause) Holttum View in CoL ; S. baramensis (C. Chr.) Holttum View in CoL ; S. bakeri (Harr.) S.E. Fawc. & A.R. Sm. View in CoL ; S. batacorum (Rosenst.) Holttum View in CoL ; * S. batjanensis (Rosenst.) Holttum View in CoL ; S. batulantensis Holttum View in CoL ; * S. beccarianus (Ces.) S.E. Fawc. & A.R. Sm. View in CoL ( Figs. 7A, 7B View FIG ); S. benoitianus (Gaudich.) Holttum; * S. braithwaitei Holttum View in CoL ; * S. brauseanus (Holttum) S.E. Fawc. & A.R. Sm. View in CoL ; S. bulusanicus (Holttum) S.E. Fawc. & A.R. Sm. View in CoL ; * S. canescens (Blume) Holttum View in CoL ; S. carrii Holttum View in CoL ; S. cartilagidens P.M. Zamora & Co View in CoL ; S.castaneus (A.R. Sm. & Lorence) A.R.Sm. & Lorence View in CoL ; * S. cataractorum (W.H.Wagner & Grether) Holttum View in CoL ; S. caulescens Holttum View in CoL ; * S. confertus (Brause) Holttum View in CoL ; S. convergens Holttum View in CoL ; S. cyrtocaulos (Alderw.) Holttum View in CoL ; S. daymanianus Holttum View in CoL ; S. debilis (Mett.) Holttum View in CoL ; * S. decadens (Baker) Holttum View in CoL ; * S. dichrotrichoides (Alderw.) Holttum View in CoL ; S. dichrotrichus (Copel.) Holttum View in CoL ; S. dimidiolobatus Holttum View in CoL ; S. diminutus (Copel.) M.G. Price View in CoL ; S. dimorphus (Brause) Holttum View in CoL ; S. diversilobus (C. Presl) Holttum View in CoL ; * S.doodioides (Copel.) Holttum View in CoL ( Fig. 2E View FIG ); S. echinosporus (Alderw.) Holttum View in CoL ; S. efogensis Holttum View in CoL ; S. ekutiensis Holttum View in CoL ; S. ellipticus (Rosenst.) Holttum View in CoL ; S. eminens (Baker) Holttum View in CoL ; S. erectus (Copel.) Holttum View in CoL ; S. erwinii Holttum View in CoL ; S. exindusiatus Holttum View in CoL ; S. flavoviridis Holttum View in CoL ; S. foliolosus Holttum View in CoL ; S. foxworthyi Holttum View in CoL ; S. gaudichaudii Holttum View in CoL ; * S. glandulosus (Blume) S.E. Fawc. & A.R. Sm. View in CoL ; S. grandescens Holttum View in CoL ; S. gregarius (Copel.) Holttum View in CoL ; S. gymnorachis Holttum View in CoL ; S. hamiferus (Alderw.) Holttum View in CoL ; * S. hastatopinnatus (Brause) Holttum View in CoL ; S. hellwigensis Holttum View in CoL ; S. hendersonii Holttum View in CoL ; S. hernaezii Holttum View in CoL ; * S. heterocarpos (Blume) Holttum ( Fig. 2K View FIG ); S. heterophyllus (C. Presl) S.E. Fawc. & A.R. Sm. View in CoL ; * S. hirsutus (Kunze ex Mett.) Holttum View in CoL ; S. hispidifolius (Alderw.) Holttum View in CoL ; S. hispiduliformis (C. Chr.) Holttum View in CoL ; S. hoalensis Holttum View in CoL ; S. hochreutineri (Christ) Holttum View in CoL ; S. humilis Holttum View in CoL ; S. immucosus Holttum View in CoL ; * S. incisus (Copel.) S.E. Fawc. & A.R. Sm. View in CoL ; S. inconspicuus (Copel.) Holttum View in CoL ; S. indrapurae Holttum View in CoL ; S. intermedius S.J. Lin View in CoL ; S. irayensis (Copel.) Holttum View in CoL ; S. isomorphus Holttum View in CoL ; S. kalkmanii Holttum View in CoL ; * S. lamii Holttum View in CoL ; * S. larutensis (Bedd.) C. Chr. View in CoL ; * S. lastreoides (C. Presl) Holttum View in CoL ; * S. latebrosus (Kunze ex Mett.) Holttum View in CoL ; S. lithophyllus (Copel.) Holttum View in CoL ; S. lobangensis (C. Chr.) Holttum View in CoL ; S. lobatus (Copel.) Holttum View in CoL ; S. loherianus (Christ) Holttum View in CoL ; S. longbawanensis (K. Iwats. & M. Kato) S.E. Fawc. & A.R. Sm. View in CoL ; * S. maemonensis (W.H. Wagner & Grether) Holttum View in CoL ; * S. magnus (Copel.) Holttum View in CoL ; S. major (Copel.) Holttum ; S. makassaricus Holttum View in CoL ; S. maximus (K. Iwats. & M. Kato) S.E. Fawc. & A.R. Sm. View in CoL ; S. melanophlebius (Copel.) S.E. Fawc. & A.R. Sm. View in CoL ; S. melanorachis Holttum View in CoL ; S. menadensis Holttum View in CoL ; S. mengenianus Holttum View in CoL ; S. metcalfei (Baker) Holttum View in CoL ; S. microlonchus (Christ) S.E. Fawc. & A.R. Sm. View in CoL ; S. micropinnatus (Holttum) S.E. Fawc. & A.R. Sm. View in CoL ; S. mindorensis Holttum View in CoL [name unaccounted for in Holttum, 1982]; S. mjobergii Holttum View in CoL ; S. morotaiensis Holttum View in CoL ; S. moseleyi Holttum View in CoL ; S. multiauriculatus (Copel.) Holttum View in CoL ; S. muluensis Holttum View in CoL ; S. mundus (Rosenst.) Holttum View in CoL ; S. mutabilis (Brause) Holttum View in CoL ; S. nakaikei Holttum View in CoL ; S. neotoppingii Holttum View in CoL ; S. norrisii (Rosenst.) Holttum View in CoL ; S. novae-britanniae Holttum View in CoL ; * S. novoguineensis (Brause) Holttum View in CoL ; S. nudisorus Holttum View in CoL ; S. obtusifolius (Rosenst.) Holttum View in CoL ; S. omatianus Holttum View in CoL ; * S. oosorus (Baker) Holttum View in CoL ; S. paripinnatus (Copel.) Holttum View in CoL ; S. penniger (Hook.) Holttum View in CoL ; * S. pentaphyllus (Rosenst.) S.E. Fawc. & A.R. Sm. View in CoL ; S. perglanduliferus (Alderw.) Holttum View in CoL ; S. petiolatus (K. Iwats. & M. Kato) S.E. Fawc. & A.R. Sm. View in CoL ; S. pilosissimus Holttum View in CoL ; S. pilosiusculis (Racib.) S.E. Fawc. & A.R. Sm. ; S. pilososquamatus (Alderw.) Holttum View in CoL ; S. plurifolius (Alderw.) Holttum View in CoL ; S. plurivenosus Holttum View in CoL ; S. polisianus Holttum View in CoL ; * S. polycarpos (Blume) Copel. ( Fig. 2D View FIG ); * S. polyotis (C. Chr.) Holttum View in CoL ; S. porphyricola (Copel.) Holttum View in CoL ; S. posthumii Holttum; S. potamios Holttum View in CoL ; * S. productus (Kaulf.) Holttum View in CoL ; S. pseudomegaphyllus (Alderw.) Holttum View in CoL ; S. pterosporus (Alderw.) Holttum View in CoL ; S. pullenii Holttum View in CoL ; S. pycnosorus (C. Chr.) Holttum View in CoL ; S. reconditus Holttum View in CoL ; S. reineckei (C. Chr.) Holttum View in CoL ; S. richardsii (Baker) Holttum View in CoL ; S. rigidus (Ridl.) Holttum View in CoL ; S. roemerianus (Rosenst.) Holttum View in CoL ; S. rudis (Ridl.) Holttum View in CoL ; S. sagittifolius (Blume) Holttum View in CoL ; S. sakayensis (Zeiller) Holttum View in CoL in B. K. Nayar & S. Kaur; S. santomasii Holttum View in CoL ; * S. sarasinorum Holttum View in CoL ; S. scandens Holttum View in CoL ; S. scopulorum (Holttum) S.E. Fawc. & A.R. Sm. View in CoL ; S. semicordatus Holttum View in CoL ; S. semimetralis Holttum View in CoL ; * S. sessilipinna (Copel.) Holttum View in CoL ; S. simplicifolius (J. Sm. ex Hook.) Holttum View in CoL ; S. solutus Holttum View in CoL ; S. spenceri (Christ) Holttum View in CoL ; S. squamatellus Holttum View in CoL ; S. stenodontus (Copel.) Holttum View in CoL ; * S. stipellatus (Blume) Hottum View in CoL ; S. stresemannii Holttum View in CoL ; S. subalpinus (Alderw.) Holttum View in CoL ; S. subappendiculatus (Copel.) S.E. Fawc. & A.R. Sm. View in CoL ; S. subcanescens Holttum View in CoL ; S. subcordatus Holttum View in CoL ; * S. suboppositus Holttum View in CoL ; * S. subpectinatus (Copel.) Holttum View in CoL ; S. subulifolius (Alderw.) Holttum View in CoL ; S. subtruncatus (Bory) Holttum View in CoL ; S. sudesticus Holttum View in CoL ; S. superbus (Brause) S.E. Fawc. & A.R. Sm. View in CoL ; * S. taiwanensis (C. Chr.) Holttum ex Kuo View in CoL ; S. tandikatensis (Alderw.) Holttum View in CoL ; S. telefominicus Holttum View in CoL ; S. tephrophyllus (Copel.) Holttum View in CoL ; S. tibangensis (C. Chr.) Holttum View in CoL ; S. trichochlamys Holttum View in CoL ; S. trimetralis Holttum View in CoL ; S. uaniensis Holttum View in CoL ; S. uniauriculatus (Copel.) Holttum View in CoL ; * S. urdanetensis (Copel.) Holttum View in CoL ; * S. veitchii Holttum View in CoL ; * S. vestigiatus (Copel.) Holttum View in CoL ; * S. warburgii (Kuhn & Christ) Holttum View in CoL ; S. wauensis Holttum View in CoL ; S.williamsii (Copel.) Holttum View in CoL ; S. woitapensis Holttum View in CoL ; S. womersleyi (Holttum) S.E. Fawc. & A.R. Sm. View in CoL

Excluded species.— Sphaerostephanos invisus (G. Forst.) Holttum and S. unitus (L.) Holttum are both combined in Strophocaulon . Sphaerostephanos cucullatus (Blume) S.M. Almeida & M.R. Almeida was treated as a synonym of S. unitus by Holttum (1982), and so presumably is a synonym of Strophocaulon unitum (L.) S. E. Fawc. & A. R. Sm., or related to that. Sphaerostephanos elatus (Bojer) Holttum is inferred as sister to Pseudocyclosorus in Fawcett et al. (in press), but resolves with species of Abacopteris in earlier studies (e.g., Patel et al. 2019a). Two species, Sphaerostephanos hirtisorus (C. Chr.) Holttum and Sphaerostephanos peltochlamys (C. Chr.) Holttum are here combined in Abacopteris .

Incertae sedis.—We were unable to place Sphaerostephanos kurzii Holttum. Sphaerostephanos validus (Christ) Holttum , is considered a synonym of Abacopteris hirtisora by Lindsay et al. (2009) but is in need of further study. Nannothelypteris seramensis M. Kato may belong in Sphaerostephanos based on its morphology, although we treat all other species of Nannothelypteris in Pronephrium .

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Abbreviations

S	Department of Botany, Swedish Museum of Natural History
B	Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet
K	Royal Botanic Gardens
E	Royal Botanic Garden Edinburgh
A	Harvard University - Arnold Arboretum
R	Departamento de Geologia, Universidad de Chile

Copyright notice

No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.

Taxonomy

Kingdom	Plantae
Phylum	Tracheophyta
Class	Polypodiopsida
Order	Polypodiales
Family	Thelypteridaceae
Genus	Sphaerostephanos

Sphaerostephanos womersleyi (Holttum) S.E. Fawc.& A.R. Sm.

Fawcett, Susan & Smith, Alan R. 2021

2021

Pronephrium womersleyi Holttum,Blumea

Holttum 1972: 108

1972

Specimens

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