Roddenberryus, Sánchez-Ruiz & Bonaldo, 2023
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publication ID |
https://doi.org/ 10.5852/ejt.2023.891.2263 |
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publication LSID |
lsid:zoobank.org:pub:F0219C1F-DFB9-4BB2-A295-4898CFF38F44 |
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DOI |
https://doi.org/10.5281/zenodo.8325478 |
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persistent identifier |
https://treatment.plazi.org/id/03B787CE-5867-FFD5-A02C-40A3FAFB67FF |
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treatment provided by |
Felipe |
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scientific name |
Roddenberryus |
| status |
gen. nov. |
Genus Roddenberryus gen. nov.
urn:lsid:zoobank.org:act:33515347-F78C-4513-8F8A-B509087E1CEF
Type species
Roddenberryus kirk gen. et sp. nov. (here designated).
Diagnosis
Members of Roddenberryus gen. nov. can be distinguished from all non-nopine genera by the presence of tarsal adesmatic joints ( Figs 1H View Fig , 5G View Fig , 9H View Fig , 12H View Fig ) and from all other Nopinae by a conformation of the unique characters of the endites and labium among these genera, in which the endites have an accentuated finger-shaped forward projection and the labium is triangular and projected ( Figs 1E View Fig , 2C View Fig , 3C View Fig , 4C View Fig , 8F View Fig , 9E View Fig , 12E View Fig , 14C View Fig ). Additionally, members of Roddenberryus gen. nov. can be distinguished by the triangular, very short, scaly gladius ( Figs 5G, H View Fig , 9H View Fig , 12H View Fig ), strongly projected tarsal organ margins ( Fig. 6E–F View Fig ) and the serrula composed by interspersed multiple rows of teeth ( Fig. 4F View Fig ).
Etymology
The generic name, masculine, is a patronymic honoring Eugene Wesley Roddenberry Sr, the creator of Star Trek, a science fiction media franchise that inspired generations of kids to pursue scientific careers.
Other species included
Roddenberryus spock gen. et sp. nov., R. mccoy gen. et sp. nov., R. sargi (F. O. Pickard-Cambridge, 1899) gen. et comb. nov. and R. pelegrina ( Bryant, 1940) gen. et comb. nov.
Description
Caponiids with only two eyes ( Fig. 1D View Fig ). Carapace orange, elongate oval, widest at rear of coxae II, only gradually narrowed anterior of eyes, without conspicuous pattern ( Figs 1D View Fig , 9D View Fig , 12D View Fig ). Pars cephalica flattened, pars thoracica slight sloping posteriorly ( Fig. 4A View Fig ); thoracic groove absent. Anterior median eyes dark, situated on slightly elevated black ocular tubercle, separated by about a half of its diameter, set back from anterior margin of clypeus by about twice their diameter ( Figs 1D View Fig , 9D View Fig , 12D View Fig ). Chelicerae orange, with median lamina; most of distance between lamina and fang base occupied by white membranous lobe ( Fig. 4G–J View Fig ); cheliceral paturon with scattered, long, weak bristles ( Fig. 4H–I View Fig ); ectal side with stridulatory ridges ( Fig. 4I View Fig ); fang very short ( Fig. 4J View Fig ). Endites orange except for anterior tips due to a white membranous projection, finger-shaped, convergent along midline, but not touching ( Figs 1E View Fig , 2C View Fig , 4C View Fig , 8F View Fig , 9E View Fig , 12E View Fig ), covered with scattered long setae, and with strong distal serrula consisting of interspersed multiple tooth rows ( Fig. 4E–F View Fig ). Labium orange, triangular, projected, with broad base, fused to sternum along obsoleted posterior groove ( Fig. 4C View Fig ). Sternum orange, oval, surface with fine reticular lines with numerous long, stiff setae ( Figs 1E View Fig , 4B View Fig , 8F View Fig , 9E View Fig , 12E View Fig ); pleural membrane with three sclerotized intercoxal extensions between coxae I and II, II and III, and III and IV ( Fig. 4A View Fig ); long and thin precoxal triangles on coxae II, III and IV ( Figs 1E View Fig , 4B View Fig , 8F View Fig , 9E View Fig , 12E View Fig ). Legs orange, formula 4123, without spines; anterior femora slightly thickened ( Figs 1I View Fig , 5A View Fig ); all metatarsi with multiple adesmatic joints intertwined on cuticle, specific limits poorly defined ( Figs 1H View Fig , 5G View Fig , 9H View Fig , 12H View Fig ), with dorsal metatarsal stopper ( Fig. 5I View Fig ), I–II with a crista occupying almost all ventral part ( Fig. 5G View Fig ) and a triangular, very short, scaly gladius ( Figs 1H View Fig , 5C, G, H View Fig , 9H View Fig , 12H View Fig ); all tarsi with three claws ( Fig. 5E View Fig ) and multiple adesmatic joints intertwined on cuticle ( Figs 1H View Fig , 5D View Fig , 9H View Fig , 12H View Fig ); paired claws usually with 8–10 teeth, most distal one largest ( Figs 5E–F View Fig , 6B View Fig ); unpaired claw short, with 2–4 teeth ( Fig. 5E, L View Fig ); ventral frictional setae on tarsi ( Fig. 5E, L View Fig ) and several other setae around pretarsal claws. Tibiae, metatarsi and tarsi with trichobothria in a single row, bases with semicircular rim bearing slight longitudinal ridges ( Fig. 6D View Fig ), tarsal organ exposed, but with marginal ring strongly pronounced ( Fig. 6E–F View Fig ). Female palpal tarsus not elongated, without claw, prolateral surfaces covered with strong, long setae ( Figs 1B–C View Fig , 9B–C View Fig , 12B–C View Fig ), with an oval pad of fine chemoreceptor setae on dorsal, distal sector ( Fig. 4K–L View Fig ), without a tibial brush on prolateral side, but palpal tibia also covered with strong, long setae on prolateral side ( Figs 1B View Fig , 4K View Fig , 9B View Fig , 12B View Fig ); prolateral pick on palpal femur, located almost in middle of podomere ( Figs 1B View Fig , 9B View Fig , 12B View Fig ). Abdomen pale gray dorsally ( Figs 1A View Fig , 2A View Fig , 3A View Fig , 8A View Fig , 9A View Fig , 12A View Fig ), lighter ventrally, with sclerotized epigastric and postepigastric scuta ( Figs 1F View Fig , 2B View Fig , 3B View Fig , 9F View Fig , 12F View Fig ); with two pairs of respiratory spiracles clustered around epigastric groove; anterior spiracles leading to short tracheal trunk ending in numerous long tracheoles; posterior spiracles leading to only one wide tracheal trunk extending anteriorly into cephalothorax and several short, small tracheoles extending posteriorly ( Fig. 11B, D, F View Fig ). Six spinnerets in typical caponiid arrangement ( Fig. 7G View Fig ), PMS with one major ampullate gland spigot and 10–11 aciniform gland spigot fields ( Fig. 7I View Fig ); PLS with several aciniform gland spigots ( Fig. 7K View Fig ); ALS with one presumed piriform gland spigot ( Fig. 7J View Fig ); PLS considerably greater than ALS ( Fig. 7G View Fig ). External female genitalia with a sclerotized anterior plate ( Fig. 11A, C, E View Fig ); posterior plate narrower, sclerotized; external sclerotization around spiracles (ess) pointy anteriorly on anterior spiracles and tear-shaped on posterior spiracles ( Fig. 11A, C, E View Fig ). Internal female genitalia consisting of a transverse, sclerotized, anteriorly directed posterior receptacle (pr) or interpulmonary fold, a presumed uterus externus (ue) and a membranous anteromedian receptacle formed by a membranous duct at base (arb) protruding anteriorly from bursa, leading to large, oval, membranous sac-like structure (sac) ( Figs 7A–D View Fig , 11B, D, F View Fig ). Male palpal patella short, with a few short setae on prolateral surface ( Fig. 8G–H View Fig ); tibia partially excavated ventrally, with prolateral surface densely covered with strong, long setae ( Figs 2D–F View Fig , 8G–H View Fig ); cymbium elongated, swollen, partially curved, with wide tip, proximal half of prolateral surface densely covered with strong, long setae ( Figs 2D View Fig , 8G View Fig ), distal half of prolateral surface and entire retrolateral surface with few short setae ( Figs 2F View Fig , 8H View Fig ), with oval dorsal distal pad of fine chemoreceptor setae; tegulum sub-spherical, with retrolateral groove which partially divides one sub-apical third ( Figs 2D–F View Fig , 8G–H View Fig ); embolus thin, pointed, curved upward on lateral position ( Figs 2D–F View Fig , 8G–H View Fig ).
Distribution
Mexico, Costa Rica, Guatemala and Cuba ( Figs 6G View Fig , 15 View Fig ).
Key to the species of Roddenberryus gen. nov.
1. Males..................................................................................................................................................2
– Females..............................................................................................................................................3
2. Embolus short, not reaching half the length of tegulum; elongated tibia, width less than half the length ( Fig. 8G–H View Fig ) ........................................................................ R. mccoy gen. et sp. nov.
– Embolus long, greater than half the length of tegulum; thickened tibia, width almost equal to length ( Fig. 2D–F View Fig ) ........................................................................................... R. kirk gen. et sp. nov.
3. Invaginations on clypeus margins absent ( Fig. 1D View Fig ) .................................... R. kirk gen. et sp. nov.
– Invaginations on clypeus margins present ( Figs 9D View Fig , 12D View Fig , 14A View Fig )...................................................4
4. A few adesmatic joints occupying only last third of tarsus ( Fig. 12H View Fig ); epigastric furrow with median invagination and rounded laterals ( Figs 11E View Fig , 12F View Fig ) ...... R. sargi ( Pickard-Cambridge, 1899)
– Several adesmatic joints occupying almost entire tarsus ( Fig. 9H View Fig ); epigastric furrow straight with slightly invagination ( Figs 9F View Fig , 11C View Fig , 14B View Fig )..............................................................................5
5. Apical sector of tibia with dense tuft of black setae ( Fig. 14A View Fig ) ......... R. pelegrina ( Bryant, 1940)
– Apical sector of tibia lacking dense tuft of black setae .......................... R. spock gen. et sp. nov.
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