Pantodinus klugi Burmeister, 1847

Seidel, Matthias, Arriaga-Varela, Emmanuel & Sousa, Rafael, 2018, Catalogue of the Incini with the description of the first Archedinus species from Honduras (Coleoptera: Scarabaeidae: Cetoniinae), Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2), pp. 389-405 : 402

Pantodinus klugi Burmeister, 1847

( Figs 5K‒M )

Pantodinus klugii Burmeister, 1847: 292 [original combination] Pantodinus klugi: CANDEZE (1873) : XLI [incorrect subsequent spelling, here conserved according to Art. 33.3.1 of ICZN (1999)]

Jeannelosis auriculatus Bourgin, 1945 . Junior subjective synonym. Jeannelosis auriculatus Bourgin, 1945: 120 [original combination].

Synonymized by HOWDEN (1972: 653).

Types. Pantodinus klugi . HOWDEN (1972) stated that the Pantodinus klugi type location is unknown to him. Burmeister described the species based on specimen(s) from the Melly collection,which is housed in MNHG.The MNHG houses one male specimen labelled “Coll. Melly. sans patrie” and “592/3 coll. Melly”. If this this specimen is a syntype remains unclear. Jeannelosis auriculatus: One male holotype in MNHN ( BOURGIN 1945). Locality records from literature. GUATEMALA: CHIMALTENANGO (MORÓN & VAZ- DE- MELLO 2007), GUATEMALA ( HOWDEN 1972), QUETZALTENANGO (MORÓN & VAZ- DE- MELLO 2007), SACATEPÉQUEZ ( HOWDEN 2010), SAN MARCOS (MORÓN & VAZ- DE- MELLO 2007), SOLOLÁ ( HOWDEN 1972, MORÓN & VAZ- DE- MELLO 2007). MEXICO: CHIAPAS ( MORÓN et al. 1997, MORÓN & VAZ- DE- MELLO 2007).

Additional material examined. GUATEMALA: 1♁ 1♀ ( RBINS): Guatemala, 15.iii.1873. CHIMALTENANGO: 3♁♁ ( DBPC): Guatemala,Zaragoza, vii.1994; 1 ♁ ( UNSM): Guatemala, Chimaltenango, San Jose Chirijuya, viii.1975; 2 ♁♁ ( BMNH): Guatelama. Rodiguez, Slope of the Volcan de Acatenango, near Patricia, 7000 ft., Godman-Salvin Coll. GUATEMALA: 5 ♁♁ 1♀ ( SLCC): Guatemala, El Aguacate, Mpio, Mixco, 1900m, iv.1978. QUETZALTENANGO: 1♁ 1♀ ( UNSM): Guatemala, Quetzaltenango, Cantel, iv.1985; 4 ♁♁ 1 ♀ ( UNSM): Guatemala, Quetzaltenango, Las Fuentes Georginas, Cerro Zunil, vi.2011, B.C. Ratcliffe leg.; 5♁♁ 2♀♀ ( BCRC): Guatemala, Quetzaltenango, Las Fuentes Georginas,Cerro Zunil, vi.2011, B. C. Ratcliffe leg. QUICHÉ: 1♁ ( USNM): Guatemala, El Quiche, Chichicastenango (7.3 km S), v.1973. SACATEPÉQUEZ: 1 ♁ ( PSPC): Guatemala: Sacatepéquez, Cerro Alux, 2260 m, 24.vi.1993, B. D.Gill leg; 1♁ ( PSPC): Guatemala, Sacatepéquez, Cerro Alux, 2260m, 9.vi,1991, B. D. Gill leg.; 4 ♁♁ ( ARPC): Guatemala, Sacatepéquez, Sn. Lucas, Cerro Alux, 2200 m, 26.vi.1996; 1 ♁ ( BCRC): Guatemala, Sacatepequez, Cerro Carmona, Antigua Guatemala, vii.2009, B. C. Ratcliffe leg.; 1 spec. ( EAPZ): Guatemala, Sacatepéquez, Cerro Alux, 12.vi.1991, H. & A. Howden leg., 013.825EAPZ; 1 ♁ 1 ♀ ( SLCC): Guatemala, San Lucas, Cerro Alux, Sacatepequez, 2200m, 25.vi.1997, Pierre Bélanger leg. SAN MARCOS: 1♀ ( ARPC): Guatemala, San Marcos, v.2009;4♁♁ 1♀ ( UNSM): Guatemala, San Marcos above La Fraternidad, vi.1997, J. Monzon leg. SOLOLÁ: 2 ♁♁ 2 ♀♀ ( SLCC): Guatemala, Solola, Xajaac, 2325 m, 15.v.1988. MEXICO: CHIAPAS: 1 ♁ ( CEMT): Mexico, Chiapas, Mpio:Angel A. Corzo, Reserva “El Triunfo”, 13.v.2005, G. Gomez leg.

Remarks. CANDEZE (1873), while transferring Pantodinus klugii Burmeister, 1847 from “Dynastides” to “Trichides”, misspelled the species epithet as “ klugi ” lacking one “ i ”. This is an incorrect subsequent spelling, which has since been used by some authors ( CANDEZE 1873; BATES 1889; KOLBE 1909; SCHENKLING 1922; BLACKWELDER 1944; HOWDEN 1968, 1972, 2010; MORÓN & KRIKKEN 1990; MORÓN et al. 1997; MORÓN & VAZ- DE- MELLO 2017). At the same time P. klugii was used only once by WESTWOOD (1878). We follow Art. 33.3.1 ( ICZN 1999), which states that “when an incorrect subsequent spelling is in prevailing usage and it is attributed to the publication of the original spelling, the subsequent spelling and attribution are to be preserved and the spelling is deemed to be a correct original spelling”. Since Pantodinus klugi Burmeister, 1847 is in prevailing usage it is deemed to be a correct original spelling. KRAJCIK (1999, 2012) omitted Jeannelosis auriculatus .

Figures

Fig 5. Habitus views of Incini species.A–C, E, G–H, J–K, M – dorsal habitus. D, F, I, L – lateral habitus. A–B – Archedinus relictus Morón & Krikken, 1990: A –♁ paratype; B –♀ paratype. C–D – Coelocratus rufipennis (Gory & Percheron, 1833), ♁ lectotype. E‒F – Inca clathratus clathratus (Olivier, 1792), ♁. G – I. clathratus sommeri Westwood, 1844, ♀. H–J – Golinca bifrons (Olivier, 1789): H–I – ♁, J –♀. K–M – Pantodinus klugi Burmeister, 1847: K–L –♁, M –♀. All scale bars = 10 mm. Images 5A–B courtesy of D. Kovac, C–D by courtesy of G. Cuccodoro, 5E‒M: by courtesy of R. Limoges.