Tonsilla subtruculenta, Zhang & Irfan & Wang & Zhang, 2022

Zhang, Meng, Irfan, Muhammad, Wang, Lu-Yu & Zhang, Zhi-Sheng, 2022, Six new species of Tonsilla Wang & Yin, 1992 (Araneae: Agelenidae) from southern China, with the first description of the male of T. yanlingensis (Zhang, Yin & Kim, 2000), Zootaxa 5091 (2), pp. 357-372 : 365-367

Tonsilla subtruculenta sp. nov.

Figures 6A–E , 9I–J , 10

Type material. Holotype ♂: CHINA, Guizhou Province, Leigong Mountain, Xian’nvtang , 26.36793333°N, 108.20136667°E, 1494 m, 21.X.2012, L.Y. Wang, X.K. Jiang & X.W. Meng leg. (SWUC-T-AG-53-01) ; Paratypes (4 ♂, 3 ♀): 2 ♂, Xian’nvtang , 26.36935°N, 108.20221667°E, 1541 m, 21.X.2012, L.Y. Wang, X.K. Jiang & X.W. Meng leg. (SWUC-T-AG-53-02~03) ; 2 ♂, Shibing County, Yuntai Mountain, 27.11061667°N, 108.12056667°E, 871 m, 18.X.2012, L.Y. Wang, X.K. Jiang & X.W. Meng leg. (SWUC-T-AG-53-04~05) ; 2 ♀, Lianhuaping , 26.361629°N, 108.164842°E, 1600 m, 14.X.2005, Z.S. Zhang & H.M. Chen leg. (SWUC-T-AG-53-06~07) ; 1 ♀, same data as holotype (SWUC-T-AG-53-08) .

Etymology. The specific epithet refers to similarity of the male to Tonsilla truculenta Wang & Yin, 1992 ; adjective.

Diagnosis. The male of this new species resembles T. truculenta ( Zhu et al. 2017) in having a similar tegular sclerite, conductor’s dorsal apophysis, tibial apophysis of male palp. The females of both species have epigynes with long epigynal teeth. Both species can be distinguished by the following characters: patellar apophysis orientated parallel to tibial base in retrolateral view in T. subtruculenta sp. nov. ( Fig. 6B, C ), vs. extending above the tibia in T. truculenta ( Zhu et al. 2017, fig. 361E). Upper part of conductor apex somewhat rectangular in ventral view in T. subtruculenta sp. nov. ( Fig. 6B ), vs. thumb-shaped in T. truculenta ( Zhu et al. 2017, fig. 361E). The female of T. subtruculenta sp. nov. can be distinguished from all congeners by the following characters: atrial ridges round with copulatory opening at the base of septum posteriorly; copulatory ducts oval, anteriorly covered by spermathecal head; spermathecae robust, situated ventro-mesally ( Fig. 6D, E ).

Description. Male (holotype, Fig. 9I ). Total length 14.87. Carapace 7.66 long, 4.71 wide; opisthosoma 7.49 long, 4.83 wide. Eye sizes and interdistances: AME 0.33, ALE 0.32, PME 0.28, PLE 0.34; AME–AME 0.13, AME– ALE 0.13, PME–PME 0.19, PME–PLE 0.35, ALE–PLE 0.08. MOA 0.88 long, front width 0.70, back width 0.84. Clypeus height 0.28. Leg measurements: I 22.20 (5.88, 7.46, 5.42, 3.44); II 20.14 (5.32, 6.50, 5.01, 3.31); III 17.52 (4.61, 5.45, 4.83, 2.63); IV 22.88 (5.97, 7.27, 6.35, 3.29). Spination of legs: femur I 220, II 400, III 600, IV 220; patella I-IV 000; tibia I-II 026, III-IV 046; metatarsus I 046, II 056, III 066, IV 067.

Palp ( Fig. 6A–C ). Patella longer than patellar apophysis; patellar apophysis almost parallel to the tibial base in retrolateral view. Retrolateral and lateral tibial apophysis distinct with blunt end. Cymbium approximately 3 times longer than wide; cymbial furrow extensively sclerotized, 1/3 of the cymbial length. Median apophysis spoon-like. Conductor apex bifurcated, upper part somewhat rectangular and lower part round. Conductor lamella distinct. Conductor’s dorsal apophysis curved clockwise. Embolus filiform, originating at the 7 o’clock position.

Female (one of the paratypes, Fig. 9J ). Total length 13.06. Carapace 6.63 long, 4.17 wide; opisthosoma 6.93 long, 4.16 wide. Eye sizes and interdistances: AME 0.18, ALE 0.27, PME 0.23, PLE 0.24; AME–AME 0.10, AME– ALE 0.12, PME–PME 0.17, PME–PLE 0.26, ALE–PLE 0.11. MOA 0.73 long, front width 0.46, back width 0.68. Clypeus height 0.22. Leg measurements: I 16.22 (4.53, 5.59, 3.64, 2.46); II 14.32 (4.19, 4.53, 3.24, 2.36); III 13.04 (3.75, 4.22, 3.14, 1.93); IV 17.53 (4.80, 5.75, 4.68, 2.30). Spination of legs: femur I-II 210, III 600, IV 400; patella I-III 010, IV 000; tibia I-II 026, III-IV 046; metatarsus I 026, II 046, III 026, IV 426.

Epigyne ( Fig. 6D, E ). Epigynal plate almost round, teeth long, reaching ventro-mesally. Epigynal hoods shallow, distinct, located ventro-mesally.Atrial ridges distinct and round.Atrium posteriorly divided with septum. Copulatory opening situated around the septum. Copulatory ducts oval. Spermathecal head extending anteriorly, curved around the copulatory ducts. Spermathecae robust, longer than wide, slightly curved, situated ventro-mesally.

Distribution. China (Guizhou) ( Fig. 10 ).

Figures

FIGURE 6. Tonsilla subtruculenta sp. nov. A. Male holotype, left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Epigyne, ventral view; E. Same, dorsal view. Abbreviations: CD, copulatory duct; CDA, Conductor’s dorsal apophysis; CF, cymbial furrow; CL, conductor lamella; Co, conductor; CO, copulatory opening; EH, epigynal hood; Em, embolus; ET, epigynal teeth; FD, fertilization ducts; LTA, lateral tibial apophysis; MA, median apophysis; PA, patellar apophysis; RTA, retrolateral tibial apophysis; SH, spermathecal heads; Sp, spermathecae.

FIGURE 9. Habitus of Tonsilla spp. A–B, male holotype and female paratype of T. jinyunensis sp. nov.; C–D, male holotype and female paratype of T. jiugongensis sp. nov.; E–F, male holotype and female paratype of T. rutunda sp. nov.; G–H, male holotype and female paratype of T. subrostrum sp. nov.; I–J, male holotype and female paratype of T. subtruculenta sp. nov.; K–L, male and female of T. yanlingensis (Zhang, Yin & Kim, 2000); M, male holotype of T. yueliangensis sp. nov.

FIGURE 10. Distribution of Tonsilla species treated here.